SCHEDULE 14A INFORMATION

                  Proxy Statement Pursuant to Section 14(a) of
            the Securities Exchange Act of 1934 (Amendment No.    )


              
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      Check the appropriate box:
      / /        Preliminary Proxy Statement
      / /        CONFIDENTIAL, FOR USE OF THE COMMISSION ONLY (AS PERMITTED
                 BY RULE 14a-6(e)(2))
      /X/        Definitive Proxy Statement
      / /        Definitive Additional Materials
      / /        Soliciting Material Pursuant to Section240.14a-12

                                         APPLERA CORPORATION
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                 (Name of Registrant as Specified In Its Charter)

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                    (Name of Person(s) Filing Proxy Statement,
                           if other than the Registrant)


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                                   [GRAPHIC]

INTERNET VOTING

    Most stockholders have a choice of voting over the Internet, by telephone,
or by using a traditional proxy card. Please check your proxy card or the
information forwarded by your bank, broker, or other holder of record to see
which options are available to you.

ELECTRONIC ACCESS TO PROXY MATERIALS AND ANNUAL REPORT

    This Notice of Annual Meeting and Proxy Statement and the 2002 Annual Report
are available on our Internet site at HTTP://WWW.APPLERA.COM. If you are a
stockholder of record and would like to view future proxy statements and annual
reports over the Internet instead of receiving copies in the mail, follow the
instructions provided when you vote over the Internet or call our toll-free
stockholder services number at 1-800-730-4001. If you hold your shares through a
bank, broker, or other holder, check the information provided by that entity for
instructions on how to elect to view future proxy statements and annual reports
and vote your shares over the Internet. Opting to receive your proxy materials
online saves us the cost of producing and mailing these materials to your home
or office and gives you an automatic link to the proxy voting site.

HOUSEHOLDING OF ANNUAL MEETING MATERIALS

    Some banks, brokers, and other nominee record holders may be participating
in the practice of "householding" proxy statements and annual reports. This
means that only one copy of this Notice of Annual Meeting and Proxy Statement
and the 2002 Annual Report may have been sent to multiple stockholders in your
household. If you would like to obtain another copy of either document, please
contact the Corporate Secretary at 301 Merritt 7, P.O. Box 5435, Norwalk, CT
06856-5435, telephone (203) 840-2000. If you want to receive separate copies of
the proxy statement and annual report in the future, or if you are receiving
multiple copies and would like to receive only one copy for your household, you
should contact your bank, broker, or other nominee record holder, or you may
contact us at the above address or telephone number.

                           DIRECTIONS TO THE MEETING

   The meeting will be held at 301 Merritt 7, Main Avenue (old U.S. Route 7),
                                    Norwalk,
   Connecticut, approximately one-half mile north of Exit 40B (northbound or
                                  southbound)
     on the Merritt Parkway (Connecticut Route 15). Signs in the lobby will
                           direct you to the meeting.

[LOGO]

301 Merritt 7
P.O. Box 5435
Norwalk, CT 06856-5435

September 4, 2002

                            NOTICE OF ANNUAL MEETING
                                OF STOCKHOLDERS

    The 2002 Annual Meeting of Stockholders of Applera Corporation will be held
at 301 Merritt 7, Main Avenue (old U.S. Route 7), Norwalk, Connecticut, on
Thursday, October 17, 2002 at 9:30 a.m. The meeting will be held for the
following purposes:

       1.  To elect ten directors;

       2.  To ratify the selection of PricewaterhouseCoopers LLP as independent
           accountants for the fiscal year ending June 30, 2003;

       3.  To approve amendments to the Applera Corporation 1999 Employee Stock
           Purchase Plan to increase the number of shares available for
           issuance;

       4.  To approve an amendment to the Applera Corporation/Applied Biosystems
           Group 1999 Stock Incentive Plan to increase the number of shares
           available for issuance;

       5.  To approve an amendment to the Applera Corporation/Celera Genomics
           Group 1999 Stock Incentive Plan to increase the number of shares
           available for issuance; and

       6.  To transact such other business as may properly come before the
           meeting or any adjournment thereof.

    Only holders of record of shares of Applera Corporation-Applied Biosystems
Group Common Stock and Applera Corporation-Celera Genomics Group Common Stock as
of the close of business on August 28, 2002 will be entitled to vote at the
meeting.

                                          By Order of the Board of Directors,
                                          Thomas P. Livingston

                                          SECRETARY

                            YOUR VOTE IS IMPORTANT!

     Whether or not you plan to attend the meeting, please vote as soon as
    possible. You may vote in person at the meeting even if you send in your
                proxy or vote over the Internet or by telephone.

                               TABLE OF CONTENTS



                                                              PAGE
                                                           

GENERAL INFORMATION.........................................    1

ELECTION OF DIRECTORS (PROPOSAL 1)..........................    2

THE BOARD OF DIRECTORS......................................    5

  Board Committees..........................................    5

  Report of the Audit/Finance Committee.....................    6

  Compensation of Directors.................................    7

OWNERSHIP OF COMPANY STOCK..................................    8

  Greater than 5% Beneficial Owners.........................    8

  Directors and Executive Officers..........................    9

  Section 16(a) Beneficial Ownership Reporting Compliance...   10

EXECUTIVE COMPENSATION......................................   11

  Report of the Management Resources Committee..............   11

  Performance Graph.........................................   15

  Summary Compensation Table................................   16

  Option Grant Tables.......................................   17

  Option Exercises and Year-End Value Tables................   19

  Retirement Benefits.......................................   19

  Employment Contracts, Termination of Employment,
    Change-in-Control, and Other Agreements.................   21

RATIFICATION OF THE SELECTION OF INDEPENDENT ACCOUNTANTS
  (PROPOSAL 2)..............................................   22

APPROVAL OF AMENDMENTS TO THE APPLERA CORPORATION 1999
  EMPLOYEE STOCK PURCHASE PLAN (PROPOSAL 3).................   23

  Summary of Proposed Amendments............................   24

  Summary of the Purchase Plan..............................   24

  Certain Federal Income Tax Consequences...................   26

  New Plan Benefits.........................................   26

  Vote Required for Approval................................   27

APPROVAL OF AMENDMENTS TO THE APPLERA CORPORATION/APPLIED
  BIOSYSTEMS GROUP 1999 STOCK INCENTIVE PLAN AND THE APPLERA
  CORPORATION/CELERA GENOMICS GROUP 1999 STOCK INCENTIVE
  PLAN (PROPOSALS 4 AND 5)..................................   27

  Summary of Proposed Amendments............................   27

  Summary of the Incentive Plans............................   28

  Certain Federal Income Tax Consequences...................   32

  New Plan Benefits.........................................   34

  Equity Compensation Plan Information......................   35

  Vote Required for Approval................................   37

OTHER BUSINESS..............................................   37

STOCKHOLDER PROPOSALS.......................................   37


[LOGO]

301 Merritt 7
P.O. Box 5435
Norwalk, CT 06856-5435

September 4, 2002

                                PROXY STATEMENT

                              GENERAL INFORMATION

    This proxy statement and the accompanying proxy card are first being sent to
stockholders on or about September 6, 2002 in connection with the solicitation
of proxies by the Board of Directors of Applera Corporation (the "Company" or
"we") for use at our 2002 Annual Meeting of Stockholders.

    THE ANNUAL MEETING.  The annual meeting will be held on Thursday,
October 17, 2002 at 9:30 a.m. at 301 Merritt 7, Main Avenue (old U.S. Route 7),
Norwalk, Connecticut. At this meeting, stockholders will be asked to elect 10
directors, ratify the selection of independent accountants for the Company, and
approve amendments to our employee stock purchase and stock incentive plans to
increase the number of shares available for issuance under these plans.

    RECORD DATE; STOCKHOLDERS ENTITLED TO VOTE.  Only holders of record of the
Company's Applied Biosystems Group Common Stock ("Applera-Applied Biosystems
stock") and the Company's Celera Genomics Group Common Stock ("Applera-Celera
Genomics stock") as of the close of business on August 28, 2002, the record date
for the meeting, are entitled to vote at the meeting. (The Applera-Applied
Biosystems stock and the Applera-Celera Genomics stock are sometimes referred to
collectively as "Applera common stock.")

    As of the record date, there were 208,792,550 shares of Applera-Applied
Biosystems stock and 71,259,060 shares of Applera-Celera Genomics stock
outstanding and entitled to vote at the meeting.

    VOTING RIGHTS.  At the meeting, each outstanding share of Applera-Applied
Biosystems stock will be entitled to one vote, and each outstanding share of
Applera-Celera Genomics stock will be entitled to 0.587 vote. The voting rights
of the Applera-Celera Genomics stock have been determined based on recent market
values of each class of Applera common stock in accordance with the formula set
forth in our Restated Certificate of Incorporation. The holders of
Applera-Applied Biosystems stock and Applera-Celera Genomics stock will vote
together as a single class at the meeting.

    QUORUM.  The presence at the meeting, either in person or by proxy, of a
majority of the total votes entitled to be cast by the outstanding shares of
Applera common stock is necessary to constitute a quorum to transact business at
the meeting. If a quorum is not present, it is expected that the meeting will be
adjourned or postponed in order to solicit additional proxies.

    Abstentions and broker non-votes are counted as present and entitled to vote
for purposes of determining a quorum. Broker non-votes occur when a broker or
other nominee does not vote on a particular proposal because the broker or other
nominee does not have discretionary voting authority for that proposal and has
not received voting instructions from the beneficial owner.

    VOTE REQUIRED.  A plurality of the votes cast at the meeting is required for
the election of directors (that is, the nominees receiving the greatest number
of votes will be elected). Accordingly, abstentions will not affect the outcome
of the election of directors.

                                       1

    The favorable vote of a majority of the votes present in person or by proxy
and entitled to vote at the meeting is required to ratify the selection of
independent accountants (Proposal 2) and to approve the amendments to our
employee stock purchase and stock incentive plans (Proposals 3, 4, and 5).
Abstentions on any of these proposals will be counted for purposes of
determining the number of shares present on the proposal but will not be counted
as votes "for" the proposal. Therefore, abstentions will have the same effect as
votes against these proposals. Broker non-votes will have no effect on these
proposals because they are not considered votes cast or shares entitled to vote
at the meeting.

    PROCEDURES FOR VOTING.  Stockholders of record (that is, stockholders who
hold shares of Applera common stock in their own names in our stock records
maintained by our transfer agent, Equiserve Trust Company, N.A.) have the choice
of voting by Internet, by telephone, or by completing and returning the
accompanying proxy card. The shares represented by a properly signed proxy card
or voted over the Internet or by telephone will be voted at the meeting as
specified by the stockholder. If a proxy card is properly signed and returned
but no specific choices are made, the shares represented by the proxy card will
be voted in favor of the election of all of the nominees for director and each
of the proposals set forth on the proxy card.

    "Street name" stockholders (that is, stockholders who hold shares of Applera
common stock through a bank, broker, or other nominee) who wish to vote at the
meeting will need to obtain a proxy form from the institution that holds their
shares and follow the voting instructions on that form.

    REVOCATION OF PROXIES.  A stockholder of record may revoke a proxy
(including a vote over the Internet or by telephone) at any time before it is
voted at the meeting by:

    - submitting a properly completed proxy with a later date;

    - voting by telephone or over the Internet at a later time;

    - filing with the Secretary of the Company a written revocation of proxy; or

    - voting in person at the meeting.

Attendance at the meeting will not of itself constitute revocation of a proxy.

    COSTS OF PROXY SOLICITATION.  We will bear the costs of soliciting proxies
for the meeting. In addition to solicitation by mail, proxies may be solicited
on behalf of the Company by directors, officers, or employees of the Company in
person or by telephone, facsimile, or other electronic means. We have retained
Morrow & Co., Inc., New York, New York, to assist in the distribution and
solicitation of proxies for a fee of $9,000, plus expenses.

    We will also reimburse brokerage houses and other custodians, nominees, and
fiduciaries holding shares of Applera common stock in their names or those of
their nominees for their expenses for sending proxy materials to the beneficial
owners of Applera common stock and obtaining their proxies.

                       PROPOSAL 1--ELECTION OF DIRECTORS

    The Board of Directors has nominated the ten persons named below for
election as directors of the Company at the meeting. Each of the nominees is
currently serving as a director of the Company and has agreed to serve if
elected. All directors are elected for a one-year term.

    If a nominee becomes unavailable to serve as a director for any reason, the
shares represented by proxies will be voted for such other person as may be
designated by the Board, unless the Board decides to leave the vacancy
temporarily unfilled or to reduce the number of directors.

    The principal occupation and certain other information about each of the
nominees as of August 26, 2002 is provided below.

    THE BOARD RECOMMENDS THAT YOU VOTE "FOR" EACH OF THE NOMINEES LISTED BELOW.

                                       2



                                                        
-----------------------------------------------------------------------------------------------------------------

                                                              Mr. Ayers is the retired Chairman and Chief
                                                              Executive Officer of The Stanley Works, a tool and
                                                              hardware manufacturer. He was an advisor to the
                                                              Chairman and Chief Executive Officer of Stanley
RICHARD H. AYERS                                              from January 1997 to October 1997 after having
59 Years Old                             [PHOTO]              served as Chairman and Chief Executive Officer of
Became Director 1988                                          Stanley from May 1989 to December 1996. Mr. Ayers
                                                              is a Trustee of MassMutual Institutional Funds and
                                                              MML Series Investment Fund and a director of
                                                              Instron Corporation.
-----------------------------------------------------------------------------------------------------------------

                                                              Mr. Belingard is President of Beaufour Ipsen Group,
                                                              a diversified French health care holding company.
                                                              He previously served as Chief Executive Officer of
                                                              bioMerieux-Pierre Fabre Group, a diversified French
JEAN-LUC BELINGARD                                            health care holding company, from 1999 to 2001 and
53 Years Old                                                  as Director General of the Diagnostics Division and
Became Director 1993                     [PHOTO]              a member of the Executive Committee of F.
                                                              Hoffmann-La Roche Ltd., a healthcare company, from
                                                              1990 to 1998. Mr. Belingard is also a director of
                                                              Laboratory Corporation of America Holdings and a
                                                              Foreign Trade Advisor to the French government.
-----------------------------------------------------------------------------------------------------------------

                                                              Dr. Hayes is the Philip Caldwell Professor of
ROBERT H. HAYES                                               Business Administration, Emeritus, at the Harvard
66 Years Old                                                  Business School. He has held various positions at
Became Director 1985                     [PHOTO]              Harvard since 1966. Dr. Hayes is also a director of
                                                              Helix Technology, Inc.
-----------------------------------------------------------------------------------------------------------------

                                                              Dr. Levine is a visiting professor at the Institute
                                                              for Advanced Studies. He previously served as
                                                              President and Chief Executive Officer of
ARNOLD J. LEVINE                                              Rockefeller University from 1998 to 2002 and was
63 Years Old                             [PHOTO]              the Harry C. Weiss Professor of the Life Sciences
Became Director 1999                                          and Chairman of the Molecular Biology Department at
                                                              Princeton University from 1984 to 1998. Dr. Levine
                                                              is also a director of ImClone Systems Incorporated.
-----------------------------------------------------------------------------------------------------------------

                                                              Mr. Martin is the retired President and Chief
                                                              Executive Officer of Barnes Group Inc., a
                                                              manufacturer of precision springs and custom metal
THEODORE E. MARTIN                                            components. He joined Barnes Group in 1990 as a
63 Years Old                             [PHOTO]              group vice president and served as President and
Became Director 1999                                          Chief Executive Officer from 1995 until his
                                                              retirement in 1998. Mr. Martin is also a director
                                                              of Ingersoll-Rand Company and Unisys Corporation.
-----------------------------------------------------------------------------------------------------------------


                                       3



                                                        
-----------------------------------------------------------------------------------------------------------------

                                                              Dr. Slayman is the Sterling Professor of Genetics
                                                              and Deputy Dean for Academic and Scientific Affairs
                                                              at Yale University School of Medicine. She joined
CAROLYN W. SLAYMAN                                            the Yale faculty in 1967. Dr. Slayman is a
65 Years Old                                                  consultant to the National Institutes of Health,
Became Director 1994                     [PHOTO]              most recently having served as a member of the
                                                              National Advisory General Medical Sciences Council,
                                                              and a member of the Board of Overseers of Dartmouth
                                                              Medical School.
-----------------------------------------------------------------------------------------------------------------

                                                              Mr. Smith is the retired Chairman and Chief
                                                              Executive Officer of Engelhard Corporation, a
                                                              provider of environmental technologies, specialty
ORIN R. SMITH                                                 chemical products, and engineered materials. He
67 Years Old                             [PHOTO]              served as Chairman and Chief Executive Officer of
Became Director 1995                                          Engelhard from 1995 until his retirement on January
                                                              1, 2001. He is also a director of Ingersoll-Rand
                                                              Company and Vulcan Materials Company.
-----------------------------------------------------------------------------------------------------------------

                                                              Mr. St. Laurent is a Principal of St. Laurent
                                                              Properties, a company engaged in various real
GEORGES C. ST. LAURENT, JR.                                   estate, agricultural, and forestry related
66 Years Old                             [PHOTO]              ventures. He previously served as Chief Executive
Became Director 1996                                          Officer of Western Bank from January 1988 to April
                                                              1997.
-----------------------------------------------------------------------------------------------------------------

                                                              Mr. Tobin has served as President and Chief
                                                              Executive Officer of Boston Scientific Corporation,
                                                              a medical device manufacturer, since March 1999.
                                                              Mr. Tobin previously served as President and Chief
                                                              Executive Officer of Biogen, Inc., a biotechnology
JAMES R. TOBIN                                                company, from 1997 to 1998 and President and Chief
58 Years Old                             [PHOTO]              Operating Officer from 1994 to 1997. Prior to
Became Director 1999                                          joining Biogen, he held various positions at Baxter
                                                              International Inc. including President and Chief
                                                              Operating Officer from 1992 to 1994. Mr. Tobin is
                                                              also a director of Boston Scientific and Curis,
                                                              Inc.
-----------------------------------------------------------------------------------------------------------------

                                                              Mr. White has served as Chairman, President and
                                                              Chief Executive Officer of the Company since
                                                              September 1995. Prior to that date, he was
                                                              Executive Vice President and a member of the Office
TONY L. WHITE                                                 of the Chief Executive of Baxter International
56 Years Old                             [PHOTO]              Inc., a manufacturer of health care products and
Became Director 1995                                          instruments. He also served as Group Vice President
                                                              of Baxter from 1986 to 1992. Mr. White is also a
                                                              director of AT&T Corp., C.R. Bard, Inc., and
                                                              Ingersoll-Rand Company.
-----------------------------------------------------------------------------------------------------------------


                                       4

                             THE BOARD OF DIRECTORS

    The business of the Company is managed under the direction of the Board of
Directors. Members of the Board are kept informed of the Company's business
through discussions with the Chairman, officers, and other employees, by
reviewing materials relating to the Company, and by participating in meetings of
the Board and its committees.

    During fiscal year 2002, the Board held seven meetings and the committees
held a total of 20 meetings. Average attendance at all meetings of the Board and
committees during the fiscal year was more than 94%, and each nominee for
director attended at least 75% of the meetings of the Board and of the
committees on which he or she served.

BOARD COMMITTEES

    The Board has established standing committees to assist it in carrying out
its responsibilities. The principal responsibilities of each of these committees
are described below.

    AUDIT/FINANCE COMMITTEE.  The Audit/Finance Committee oversees accounting,
finance, and internal control matters. The committee is responsible for the
appointment, compensation, and oversight of the work of the independent
accountants for the purpose of preparing or issuing an audit report or related
work. In addition, the committee: reviews and approves the scope and plan of
audit and non-audit services to be performed by the independent accountants;
reviews with management and independent accountants the Company's annual and
quarterly consolidated financial statements; reviews internal accounting and
auditing procedures; reviews the Company's financial policies and strategies;
and reviews policies and practices designed to assure the Company's compliance
with legal and ethical standards. The Audit/Finance Committee met eight times
during fiscal year 2002. A report of the committee for fiscal year 2002 is
provided on page 6.

    The current members of the Audit/Finance Committee are Richard H. Ayers
(Co-Chair), Theodore E. Martin (Co-Chair), Robert H. Hayes, and James R. Tobin.

    EXECUTIVE COMMITTEE.  The Executive Committee has the authority during the
intervals between meetings of the Board to exercise the powers of the Board
(except for certain powers reserved solely for the Board) in situations,
generally arising from unforeseen events, necessitating Board action before a
meeting can be convened. The Executive Committee did not meet during fiscal year
2002.

    The current members of the Executive Committee are Tony L. White (Chair),
Richard H. Ayers, Robert H. Hayes, and Carolyn W. Slayman.

    MANAGEMENT RESOURCES COMMITTEE.  The Management Resources Committee is
comprised of non-employee directors. The committee reviews and approves all
forms of remuneration for the senior management of the Company and administers
the Company's stock plans. It also reviews management development and succession
programs. The Management Resources Committee met eight times during fiscal year
2002. The Committee's report on executive compensation starts on page 11.

    The current members of the Management Resources Committee are Orin R. Smith
(Co-Chair), Georges C. St. Laurent, Jr. (Co-Chair), Jean-Luc Belingard, Arnold
J. Levine (effective August 15, 2002), and Carolyn W. Slayman.

    NOMINATING/CORPORATE GOVERNANCE COMMITTEE.  The Nominating/Corporate
Governance Committee (formerly designated as the Nominating Committee):
recommends nominees to fill vacancies on the Board; reviews the functioning and
effectiveness of the Board, its committees, and its individual members; and
recommends the compensation of non-employee directors and membership assignments
for committees of the Board. The Nominating/Corporate Governance Committee met
three times during fiscal year 2002.

                                       5

    The committee will consider responsible recommendations by stockholders of
candidates to be nominated as directors of the Company. All recommendations must
be in writing and addressed to the Secretary of the Company and satisfy any
applicable requirements under the Company's By-laws.

    The current members of the Nominating/Corporate Governance Committee are
Jean-Luc Belingard (Co-Chair), Carolyn W. Slayman (Co-Chair), Arnold J. Levine
(effective August 15, 2002), Orin R. Smith, and Georges C. St. Laurent, Jr.

    TECHNOLOGY ADVISORY COMMITTEE.  The Technology Advisory Committee advises
the Board and management concerning issues related to the development and
implementation of the Company's technological assets, including strategies for
developing and expanding these assets and assisting management in assessing
third party technology opportunities. The Technology Advisory Committee met one
time during fiscal year 2002.

    The current members of the Technology Advisory Committee are Arnold J.
Levine (Co-Chair), Carolyn W. Slayman (Co-Chair), Jean-Luc Belingard, Robert H.
Hayes, and Georges C. St. Laurent, Jr.

REPORT OF THE AUDIT/FINANCE COMMITTEE

    The Audit/Finance Committee assists the Board in its oversight of the
quality and integrity of the accounting, auditing, and financial reporting
practices of the Company. The Committee operates under a written charter adopted
by the Board. The Committee is comprised of four non-employee directors, all of
whom are "independent" as defined by the rules of the New York Stock Exchange.

    In performing its oversight responsibilities, the Committee reviewed and
discussed the audited consolidated financial statements of the Company as of and
for the fiscal year ended June 30, 2002 with management and
PricewaterhouseCoopers LLP ("PwC"), the Company's independent accountants.
Management has the primary responsibility for the financial statements and the
reporting process. PwC is responsible for expressing an opinion as to whether
these financial statements are presented fairly, in all material respects, in
conformity with accounting principles generally accepted in the United States.

    The Committee also discussed with PwC their judgment as to the quality, not
just the acceptability, of the Company's accounting principles and such other
matters as are required by generally accepted auditing standards, including
those described in Statement on Auditing Standards No. 61, "COMMUNICATION WITH
AUDIT COMMITTEES."

    The Committee has received the written disclosures and the letter from PwC
required by Independence Standards Board Standard No. 1, "INDEPENDENCE
DISCUSSIONS WITH AUDIT COMMITTEES," and has discussed with PwC their
independence from the Company and its management. The Committee considered
whether the provision of non-audit services by PwC to the Company is compatible
with maintaining the independence of PwC and concluded that the independence of
PwC is not compromised by the provision of such services.

    Based on the reviews and discussions referred to above, the Committee
recommended to the Board, and the Board has approved, the inclusion of the
audited consolidated financial statements of the Company in its Annual Report on
Form 10-K for the fiscal year ended June 30, 2002, for filing with the
Securities and Exchange Commission.

    The Committee has also appointed PwC to audit the Company's consolidated
financial statements for the fiscal year ending June 30, 2003, subject to
stockholder ratification of that appointment.

                                          Audit/Finance Committee
                                          Richard H. Ayers, Co-Chair
                                          Theodore E. Martin, Co-Chair
                                          Robert H. Hayes
                                          James R. Tobin

                                          August 15, 2002

                                       6

COMPENSATION OF DIRECTORS

    ANNUAL RETAINER.  Each non-employee director receives an annual retainer of
$40,000. No additional amounts are paid for participation on committees. All
directors are reimbursed for expenses incurred in attending Board and committee
meetings.

    Each non-employee director of the Company is required to apply at least 50%
of his or her annual retainer to the purchase of Applera-Applied Biosystems
stock and Applera-Celera Genomics stock. Purchases of stock are made quarterly,
and the number of shares of each class of stock purchased is based on the ratio
of the number of shares of each class outstanding on the purchase date. The
purchase price is the fair market value of a share of the applicable class of
stock on the purchase date.

    SPECIAL ASSIGNMENTS.  Recognizing that there may be occasions where the
Company is best served by non-employee directors spending additional time in
their capacities as directors, the Board has determined that it may be
appropriate to provide additional compensation for such services. Specifically,
where a non-employee director is called upon to spend materially more time than
would ordinarily be expected, a per diem payment, not to exceed reasonable and
customary rates, will be determined and approved by the Nominating/Corporate
Governance Committee. No non-employee director received any such additional
compensation during fiscal 2002.

    STOCK OPTIONS.  Each non-employee director receives an annual grant of stock
options to purchase 8,000 shares of Applera-Applied Biosystems stock and 2,000
shares of Applera-Celera Genomics stock upon his or her election or reelection
to the Board. All options granted have an exercise price equal to the fair
market value of a share of the applicable class of stock on the date of grant,
are generally exercisable in four equal annual installments, and have a term of
ten years.

    RESTRICTED STOCK.  Each non-employee director receives a restricted stock
award of up to 1,200 shares of Applera-Applied Biosystems stock and up to 300
shares of Applera-Celera Genomics stock on the date of election or reelection to
the Board. Directors elected other than at an annual meeting are granted a pro
rata portion of such shares. Restricted stock awards vest on the date
immediately preceding the annual meeting next following the date of grant and
will be forfeited, subject to certain exceptions, if the director ceases to
serve as a member of the Board prior to that date. Prior to vesting, the
director has the right to receive cash dividends and to vote but may not
transfer or otherwise dispose of the shares.

    In addition to the share limits noted above, the aggregate dollar value of
the restricted stock grants to each non-employee director was limited to
$148,500 for the fiscal year ended June 30, 2002, and may not exceed $163,350
for the fiscal year ending June 30, 2003, increasing by 10% each year
thereafter. The aggregate dollar value is calculated by multiplying the number
of shares subject to the grants by the fair market value of the applicable class
of stock on the date of grant. To the extent that the aggregate dollar value of
such awards exceeds this dollar limit, shares of each class of stock will be
allocated based on the ratio of the number of shares of each class of stock
outstanding on the date of grant. This dollar limitation did not result in any
reduction in the number of shares of restricted stock awarded to non-employee
directors in fiscal year 2002.

    DEFERRALS.  Directors may elect to defer receipt of the cash or stock
portion of their annual retainer and their restricted stock award. The stock
portion is credited to the account of a director in units quarterly, each unit
representing one share of the applicable class of stock. Directors do not have
voting rights with respect to these units. The stock portion of a director's
account is adjusted to take into account dividends paid on the stock, and the
cash portion of a director's account is credited quarterly with interest at the
prevailing prime rate of Citibank, N.A.

    STOCK OWNERSHIP POLICY.  In order to encourage non-employee directors to
retain a financial investment in the Company, the Nominating/Corporate
Governance Committee has established a requirement that each non-employee
director retain a personal investment in Applera common stock or stock
equivalents equal to four times the annual retainer. Stock equivalents that
qualify under the policy

                                       7

include deferred shares, restricted stock subject to time-based vesting, and
vested, in-the-money stock options. Non-employee directors have a period of four
years from the date of initial election to achieve this ownership level. As of
June 30, 2002, all of the non-employee directors except Mr. Tobin, who joined
the Board in August 1999, had satisfied their individual investment goals.

    MISCELLANEOUS.  As part of the Company's overall program to promote
charitable giving, the Board has established a Director's Charitable Award
Program. Under the Program, following the death of a participating director, the
Company will donate $1,000,000 to the educational or charitable organizations
selected by the director and approved by the Company. In order to fund the
donations, the Company has acquired joint life insurance contracts on the lives
of participating directors. Each policy will insure two directors with the death
benefit payable on the death of the second director. Individual directors will
derive no financial benefit from the Program since all insurance proceeds accrue
solely to the Company.

    Non-employee directors are provided business travel accident insurance when
traveling on behalf of the Company. Directors are also eligible to participate
in the Company's matching gifts program on the same basis as the Company's
employees.

    Employee directors receive no additional compensation for service on the
Board or its committees.

                           OWNERSHIP OF COMPANY STOCK

GREATER THAN 5% BENEFICIAL OWNERS

    The following are the only persons known by the Company to own beneficially
more than 5% of the outstanding shares of either class of Applera common stock
as of August 26, 2002.



                                              AMOUNT AND NATURE                    AMOUNT AND NATURE
                                                OF BENEFICIAL                        OF BENEFICIAL
                                                OWNERSHIP OF         PERCENT         OWNERSHIP OF         PERCENT
NAME AND ADDRESS OF                            APPLERA-APPLIED          OF          APPLERA-CELERA           OF
BENEFICIAL OWNER                              BIOSYSTEMS STOCK        CLASS         GENOMICS STOCK         CLASS
-------------------                        -----------------------   --------   -----------------------   --------
                                                                                              
Capital Group International, Inc. .......          25,255,180(1)       12.1
  11100 Santa Monica Blvd.
  Los Angeles, CA 90025

Capital Research and Management Company..          31,431,500(2)       15.1
  333 South Hope Street
  Los Angeles, CA 90071

FMR Corp. ...............................                                               7,194,596(3)        10.1
  82 Devonshire Street
  Boston, MA 02109

Royce & Associates, LLC. ................                                               3,853,400(4)         5.4
  1414 Avenue of the Americas
  New York, NY 10019

Wellington Management Company, LLP.......                                               9,106,717(5)        12.8
  72 State Street
  Boston, MA 02109


------------------------------

(1)   Based on an amendment to a Schedule 13G dated May 9, 2002 filed with the
     Securities and Exchange Commission (the "SEC") by Capital Group
    International, Inc. on behalf of itself and certain affiliates, Capital
    Group International, Inc. has sole voting power with respect to 20,543,360
    shares and sole dispositive power with respect to 25,255,180 shares. Capital
    Group International disclaims beneficial ownership of all such shares.

(2)   Based on a Form 13F for the quarter ended June 30, 2002 filed with the
     SEC, Capital Research and Management Company has shared investment
    discretion (as defined) and no voting authority with respect to all of these
    shares.

(3)   Based on a Form 13F for the quarter ended June 30, 2002 filed with the
     SEC, FMR Corp. has shared investment discretion (as defined) with respect
    to all of these shares, sole voting authority with respect to 142,548
    shares, and no voting authority with respect to 7,052,048 shares.

(4)   Based on a Form 13F for the quarter ended June 30, 2002 filed with the
     SEC, Royce & Associates has sole investment discretion and sole voting
    authority with respect to all of these shares.

(5)   Based on a Form 13F for the quarter ended June 30, 2002 filed with the
     SEC, Wellington Management Company, LLP has sole investment discretion with
    respect to 8,230,756 shares, shared investment discretion (as defined) with
    respect to 875,961 shares, sole voting authority with respect to 6,314,241
    shares, shared voting authority with respect to 839,417 shares, and no
    voting authority with respect to 1,953,059 shares.

                                       8

DIRECTORS AND EXECUTIVE OFFICERS

    The following table sets forth, as of August 26, 2002, information
concerning the beneficial ownership of each class of stock by (1) all directors,
(2) each of the persons named in the Summary Compensation Table below under
"EXECUTIVE COMPENSATION," and (3) all directors and executive officers of the
Company as a group. None of these persons, other than Mr. White, beneficially
owned more than one percent of the outstanding shares of either class of Applera
common stock. Mr. White beneficially owned approximately 1.1% of the outstanding
shares of Applera-Applied Biosystems stock and 1.3% of the outstanding shares of
Applera-Celera Genomics stock. All directors and executive officers as a group
beneficially owned approximately 2.6% of the outstanding shares of
Applera-Applied Biosystems stock and approximately 4.1% of the outstanding
shares of Applera-Celera Genomics stock. Except as otherwise noted, voting and
investment power is exercised solely by the beneficial owner or is shared by the
owner with his or her spouse.



                                                                                                         NUMBER OF
                                                                               NUMBER OF SHARES          DEFERRED
                                                                                 BENEFICIALLY       SHARES BENEFICIALLY
                                                   TITLE OF CLASS                 OWNED(1,2)             OWNED(3)        TOTAL
                                                   --------------            ---------------------  -------------------  -----
                                                                                                             
NON-EMPLOYEE DIRECTORS
Richard H. Ayers........................  Applera-Applied Biosystems stock          40,844                13,757         54,601
                                          Applera-Celera Genomics stock             51,696                 3,386         55,082
Jean-Luc Belingard......................  Applera-Applied Biosystems stock          43,417                     0         43,417
                                          Applera-Celera Genomics stock             52,289                     0         52,289
Robert H. Hayes.........................  Applera-Applied Biosystems stock          29,500                17,450         46,950
                                          Applera-Celera Genomics stock             48,731                 4,393         53,124
Arnold J. Levine........................  Applera-Applied Biosystems stock          16,700                 7,486         24,186
                                          Applera-Celera Genomics stock             55,871                 1,987         57,858
Theodore E. Martin......................  Applera-Applied Biosystems stock          15,500                 6,900         22,400
                                          Applera-Celera Genomics stock             45,231                 1,843         47,074
Carolyn W. Slayman......................  Applera-Applied Biosystems stock          36,574                 5,966         42,540
                                          Applera-Celera Genomics stock             50,547                 1,505         52,052
Orin R. Smith...........................  Applera-Applied Biosystems stock          29,800                14,015         43,815
                                          Applera-Celera Genomics stock             47,731                 3,594         51,325
Georges C. St. Laurent, Jr..............  Applera-Applied Biosystems stock          49,500                     0         49,500
                                          Applera-Celera Genomics stock             53,858                     0         53,858
James R. Tobin..........................  Applera-Applied Biosystems stock          15,500                 6,093         21,593
                                          Applera-Celera Genomics stock             34,891                 1,644         36,535




                                                                                                 NUMBER OF SHARES
                                                                   TITLE OF CLASS            BENEFICIALLY OWNED(4,5)
                                                                   --------------            ------------------------
                                                                                       
NAMED EXECUTIVE OFFICERS
Tony L. White...........................................  Applera-Applied Biosystems stock            2,218,924
                                                          Applera-Celera Genomics stock                 939,394
Michael W. Hunkapiller..................................  Applera-Applied Biosystems stock            1,288,011(6)
                                                          Applera-Celera Genomics stock                 642,247
Kathy P. Ordonez........................................  Applera-Applied Biosystems stock               31,250
                                                          Applera-Celera Genomics stock                  52,825
William B. Sawch........................................  Applera-Applied Biosystems stock              592,931
                                                          Applera-Celera Genomics stock                 331,186
Dennis L. Winger........................................  Applera-Applied Biosystems stock              688,797
                                                          Applera-Celera Genomics stock                 354,297
ALL DIRECTORS AND EXECUTIVE OFFICERS AS A GROUP
  (24 PERSONS)(7).......................................  Applera-Applied Biosystems stock            5,629,759(6)
                                                          Applera-Celera Genomics stock               3,020,945


                                                   (FOOTNOTES ON FOLLOWING PAGE)

                                       9

(FOOTNOTES FOR PRECEDING PAGE)

------------------------------

(1)   Includes the following number of shares which the following have the right
     to acquire within 60 days through the exercise of vested stock options:



                                Applera-Applied Biosystems stock     Applera-Celera Genomics stock
                               ----------------------------------   -------------------------------
                                                              
Richard H. Ayers.............                27,500                             48,231
Jean-Luc Belingard...........                27,500                             48,231
Robert H. Hayes..............                23,500                             47,231
Arnold J. Levine.............                15,500                             55,571
Theodore E. Martin...........                15,500                             45,231
Carolyn W. Slayman...........                27,500                             48,231
Orin R. Smith................                25,800                             47,731
Georges C. St. Laurent,
  Jr.........................                27,500                             48,231
James R. Tobin...............                15,500                             34,891


    No voting or investment power exists with respect to these shares prior to
exercise of the stock options.

(2)   Includes for each of Messrs. Ayers, Belingard, and St. Laurent 1,200
     restricted shares of Applera-Applied Biosystems stock and 300 restricted
    shares of Applera-Celera Genomics stock, as to which the holder has sole
    voting but no investment power prior to the lapse of restrictions.

(3)   Consists of units representing full shares of stock (including restricted
     stock awards) deferred by non-employee directors (see "COMPENSATION OF
    DIRECTORS," above). No voting power exists with respect to any deferred
    share units.

(4)   Includes the following number of shares which the following have the right
     to acquire within 60 days through the exercise of vested stock options:



                               Applera-Applied Biosystems Stock     Applera-Celera Genomics Stock
                              ----------------------------------   -------------------------------
                                                             
Tony L. White...............              1,731,993                            816,662
Michael W. Hunkapiller......                877,136                            508,352
Kathy P. Ordonez............                 31,250                             12,825
William B. Sawch............                523,955                            317,830
Dennis L. Winger............                631,273                            338,925


    No voting or investment power exists with respect to these shares prior to
exercise of the stock options.

(5)   Includes 72,000 restricted shares of Applera-Applied Biosystems stock and
     18,000 restricted shares of Applera-Celera Genomics stock held by
    Mr. White, 40,000 restricted shares of Applera-Celera Genomics stock held by
    Ms. Ordonez, and 12,000 restricted shares of Applera-Applied Biosystems
    stock and 3,000 restricted shares of Applera-Celera Genomics stock held by
    Mr. Winger. Prior to vesting, Messrs. White and Winger and Ms. Ordonez have
    the right to receive dividends, if any, on and to vote, but may not sell or
    otherwise dispose of, these shares.

(6)   Includes 650 shares held by Dr. Hunkapiller's son.

(7)   Includes 4,492,056 shares of Applera-Applied Biosystems stock and
     2,620,419 shares of Applera-Celera Genomics stock which all directors and
    executive officers as a group have the right to acquire within 60 days
    through the exercise of vested stock options. No voting or investment power
    exists with respect to these shares prior to exercise of the stock options.

SECTION 16(A) BENEFICIAL OWNERSHIP REPORTING COMPLIANCE

    We are required to identify any officer, director, or beneficial owner of
more than 10% of either class of Applera common stock who failed to timely file
with the SEC and the New York Stock Exchange a required report relating to
beneficial ownership of stock under Section 16(a) of the Securities Exchange Act
of 1934. Based solely on a review of information provided to us, all persons
subject to these reporting requirements filed the required reports on a timely
basis for fiscal year 2002.

                                       10

                             EXECUTIVE COMPENSATION

REPORT OF THE MANAGEMENT RESOURCES COMMITTEE

    The Management Resources Committee (the "MRC") of the Board of Directors was
comprised of four non-employee directors during fiscal year 2002. One of the
duties of the MRC is to review and approve all forms of remuneration for the
senior management of the Company.

    OVERVIEW AND PHILOSOPHY.  The overall objectives of the Company's executive
compensation plans are to:

    - Attract and retain the highest quality talent to lead the Company;

    - Reward key executives based on business performance;

    - Provide incentives designed to maximize stockholder value; and

    - Assure that objectives for corporate and individual performance are
      established and measured.

    The Company's general compensation philosophy is that total cash
compensation should vary with the performance of the Company in attaining
financial and non-financial objectives and that any long-term incentive
compensation should be closely aligned with the interests of the stockholders.
The Company has several performance-based compensation programs in which most of
the Company's employees are eligible to participate.

    Total compensation for the majority of the Company's employees, including
its executive officers, consists of the following components:

    - Base salary;

    - Annual incentive compensation provided through a cash bonus based on
      Company and individual performance; and

    - Long-term incentive compensation provided through (1) a broad based stock
      option plan and (2) a performance unit bonus plan and restricted stock
      plan for selected executives.

    The MRC's intention is to provide a competitive total compensation package
to senior management. Competitiveness is determined based upon professionally
compiled studies of the Company's peer group and other comparable companies. The
MRC particularly focuses on competitive compensation practices for companies
engaged in biotechnology product development and manufacturing and life
sciences.

    BASE SALARY.  Each year, the MRC is provided a study of compensation trends
and practices from an independent compensation consultant in order to determine
the competitiveness of the pay structure for its senior managers. Within the
broad comparative group of companies that the consultant surveys, the MRC has
identified a group of companies which compete in similar markets and which
approximate the size of the Company in terms of employees and revenue. These
companies include five of the companies contained in the industry index selected
by the Company for purposes of the Performance Graph set forth under that
heading below.

    During the fiscal year ended June 30, 2002, Mr. White's base annual salary
was $1,000,000. The MRC established Mr. White's salary based on the Company's
overall performance during the 2001 fiscal year and competitive pay practices
relative to peer companies.

                                       11

    ANNUAL INCENTIVE COMPENSATION.  Most employees of the Company not
compensated on a commission basis participate in the Company's Incentive
Compensation Program. The MRC uses specific performance objectives for each
business unit as a basis on which to measure the performance of the Company's
employees, including its executive officers, under this program. For the Applied
Biosystems group, the MRC uses EBIT (earnings before interest and taxes) and
after-tax operating cash flow targets. These financial measures are well
recognized throughout the investment community, and the MRC believes that
achieving financial goals based upon these measures should help maximize
stockholder return. For the Celera Genomics group and Celera Diagnostics, a
joint venture between the Applied Biosystems group and the Celera Genomics
group, the MRC considers financial measures including revenue growth and
operating income as well as specific business milestones. For corporate
employees, including Mr. White and certain other executive officers, the MRC
uses a combination of the performance results for the Applied Biosystems group
and the Celera Genomics group.

    The MRC uses survey information from comparable companies in reviewing and
approving annual incentive plan participation and targets for each executive
officer. In determining annual incentive compensation awards for each executive
officer, the MRC also considers other business actions taken during the fiscal
year that contribute to the strategic growth and competitiveness of the Company.
Additionally, Mr. White, based on his review of the performance of each
executive officer (other than himself) throughout the year, may propose
modifications to reflect each officer's personal performance. These
modifications may result in an incentive compensation recommendation between 0
and 150% of target. The MRC is responsible for final approval of all incentive
compensation awards for executive officers.

    Mr. White's incentive compensation formula is based entirely on the
achievement of the Company's business and financial goals. If all corporate
goals are achieved, Mr. White would receive a minimum incentive payment of 115%
of base annual salary. For the fiscal year ended June 30, 2002, Mr. White earned
an incentive compensation award of $1,150,000. This award reflects the Company's
financial performance during fiscal year 2002 and Mr. White's continued
leadership and initiative during the year, including with respect to the
continued development of Celera Diagnostics and the transformation of the focus
of the Celera Genomics group from information to therapeutic discovery and
development.

    RESTRICTED STOCK.  In fiscal year 2002, the MRC granted restricted stock
awards to selected members of senior management in connection with key strategic
initiatives at the Celera Genomics group and Celera Diagnostics. These awards,
which are subject to time-based vesting over a period of three years, serve to
recognize a significant contribution to the development of these two business
units and to retain and motivate senior management during a period of
transition. Mr. White did not receive an award of restricted stock during fiscal
year 2002.

    PERFORMANCE UNITS.  During fiscal year 2002, the MRC made a series of grants
to members of senior management under the Company's Performance Unit Bonus Plan.
The Plan utilizes stock options and a performance unit bonus pool to convey the
value targeted by a traditional restricted stock program. Performance units
granted to executive officers under the Plan in fiscal year 2002 vest upon the
price of a share of Applera-Applied Biosystems stock or Applera-Celera Genomics
stock, as the case may be, attaining and maintaining for specified periods
certain price targets and are payable in a minimum of eight quarterly payments,
subject to continued employment and program expense limitations.

    In August 2001, the MRC granted awards under the Performance Unit Bonus Plan
to members of senior management of the Applied Biosystems group and certain
executive officers, including Mr. White. These awards consisted of two grants of
performance units and stock options. The first grant of performance units
provides for vesting in two parts upon a share of Applera-Applied Biosystems
stock attaining and maintaining price targets of $30.00 and $35.00 or more for a
period of 90 days. The

                                       12

second grant of performance units provides for vesting in two parts upon a share
of Applera-Applied Biosystems stock attaining and maintaining price targets of
$40.00 and $45.00 or more for a period of 90 days. The performance units will be
forfeited to the extent the stock price targets are not attained by August 16,
2011. Upon vesting, the holder of the performance units becomes entitled to a
pre-tax cash or Applera-Applied Biosystems stock payment of $25.00 for each
vested unit. This grant is subject to a maximum aggregate quarterly payout to
all participants of $750,000. The first option grant vests three years from the
date of grant, and the second option grant vests on the earlier of (1) five
years from the date of grant or (2) two years after the stock price targets
applicable to the second grant of performance units have been attained.

    In connection with the August 2001 grant, Mr. White was granted a total of
80,000 performance units and an equal number of options to purchase shares of
Applied Biosystems stock at an exercise price of $25.00 per share.

    STOCK OPTIONS.  The MRC believes that in order to achieve the Company's
long-term growth objectives and to align employee and stockholder interests it
is in the Company's best interest to grant stock options to both management and
non-management employees. The number of stock options granted to each employee
depends on the employee's level in the Company and the potential impact of his
or her position on the overall success of the Company.

    The MRC approves the number of options granted to each participant in the
stock incentive plans during each fiscal year. Employees may be granted options
for Applera-Applied Biosystems stock or Applera-Celera Genomics stock or both
stocks. Certain officers and employees who have responsibilities involving both
the Applied Biosystems group and the Celera Genomics group will be granted
awards in both stocks in a manner that reflects their job responsibilities. The
MRC believes that granting participants awards tied to the performance of the
group in which the participants work and, in certain cases, the other group, is
in the best interest of the Company and its stockholders.

    The exercise price of each option granted is generally the fair market value
of a share of the applicable class of stock on the date of grant. Stock options
granted in fiscal year 2002 generally vest in equal installments over a period
of four years. The MRC believes that the four year vesting period serves to
promote the retention of key employees. In general, options granted under the
Company's stock incentive plans are exercisable for a period of ten years from
the date of grant.

    Mr. White was granted options to purchase 255,000 shares of Applera-Applied
Biosystems stock at an exercise price of $20.85 per share and 85,000 shares of
Applera-Celera Genomics stock at an exercise price of $18.895 per share in
April 2002 in connection with the Company's customary annual grant of options to
employees.

    STOCK OWNERSHIP.  In order to reinforce the linkage of an executive's
financial gain with stockholder results, the MRC has established a requirement
that each executive officer of the Company retain an investment in Applera
common stock or stock equivalents equaling between one and five times the
individual's annual base salary (depending upon the individual's management
level). Mr. White is required to retain a personal investment equal to five
times his annual base salary. Executives are given a period of five years to
achieve these levels. As of June 30, 2002, all of the Named Executive Officers,
except Ms. Ordonez, had satisfied their individual investment goals.

    In addition to encouraging stock ownership by granting stock options, the
Company further encourages its employees to own Applera common stock through a
tax-qualified employee stock purchase plan, which is generally available to all
domestic and certain foreign employees. This plan allows participants to buy
both classes of Applera common stock with up to 10% of their salary (subject to
certain limits).

                                       13

    The MRC monitors on an annual basis the ownership of shares of
Applera-Applied Biosystems stock and Applera-Celera Genomics stock by senior
officers as well as their option holdings and other benefits so that their
interests are not misaligned with the two classes of Applera common stock and
with their duty to act in the best interests of the Company and its stockholders
as a whole.

    CONCLUSION.  The Company has designed its executive compensation plans, as
described above, to link the compensation of senior management with the
achievement of corporate and individual performance goals. These goals have been
established at levels that the MRC believes necessary to achieve above average
performance within the Company's industry.

    The MRC intends to continue its policy of linking executive compensation
with corporate and group performance and stockholder returns to the extent
possible through the measurement procedures described in this report.
Section 162(m) of the Internal Revenue Code generally limits the tax
deductibility of certain compensation in excess of one million dollars paid to a
company's chief executive officer and the four other most highly compensated
executives. While the Company generally seeks to maximize the deductibility of
compensation paid to its executive officers, it will maintain flexibility to
take actions that may be based on considerations other than tax deductibility.

                                          Management Resources Committee
                                          Orin R. Smith, Co-Chair
                                          Georges C. St. Laurent, Jr., Co-Chair
                                          Jean-Luc Belingard
                                          Carolyn W. Slayman

                                          August 15, 2002

                                       14

PERFORMANCE GRAPH

    The following graph compares the yearly change in the Company's cumulative
total stockholder return for the last five fiscal years with the cumulative
total return on the Standard & Poor's 500 Stock Index (the "S&P 500") and the
Dow Jones Biotechnology Group Index (the "DJ BTC"), a published industry index
that includes Applera-Applied Biosystems stock and Applera-Celera Genomics
stock.

    Cumulative total returns are calculated assuming that $100 was invested on
the last trading day of fiscal year 1997 in each of the Applera common stock,
the S&P 500, and the DJ BTC, and that all dividends were reinvested. On May 6,
1999, each share of common stock of The Perkin-Elmer Corporation, the
predecessor to the Company, was converted into one share of Applera-Applied
Biosystems stock and one-half share of Applera-Celera Genomics stock. As a
result, the graph reflects a composite return for the two new classes of Applera
common stock after that date.

                              APPLERA CORPORATION
                    Comparison of 5 Year Cumulative Returns

EDGAR REPRESENTATION OF DATA POINTS USED IN PRINTED GRAPHIC



      APPLERA CORPORATION  S&P 500  DJ BTC
                           
1997                  100      100     100
1998                79.91   130.16  127.75
1999               167.97   159.78  159.12
2000               463.27   171.36  314.08
2001               191.53   145.95   258.5
2002                118.1    119.7   152.1


                                       15

SUMMARY COMPENSATION TABLE

    The following table sets forth compensation provided to the Company's Chief
Executive Officer and the four other most highly paid executive officers of the
Company (the "Named Executive Officers").



                                                                                LONG-TERM COMPENSATION
                                                     ANNUAL                ---------------------------------
                                                  COMPENSATION                     AWARDS           PAYOUTS
                                        ---------------------------------  ----------------------  ---------
                                                                OTHER
                                                               ANNUAL       RESTRICTED     STOCK     LTIP      ALL OTHER
                                FISCAL  SALARY     BONUS       COMPEN-         STOCK      OPTIONS   PAYOUTS   COMPENSATION
NAME AND PRINCIPAL POSITION      YEAR     ($)       ($)     SATION ($)(1)  AWARDS ($)(2)  (#)(3)      ($)        ($)(4)
---------------------------     ------  -------  ---------  -------------  -------------  -------  ---------  ------------
                                                                                      
Tony L. White.................   2002   984,615  1,150,000      192,044              0    420,000          0      155,692
Chairman, President and          2001   889,227    646,834      170,014     24,094,125    337,500          0       28,285
Chief Executive Officer          2000   812,693  1,364,769      249,842              0    712,600          0   14,392,277

Michael W. Hunkapiller........   2002   529,307    462,977        2,125              0    175,000          0       78,805
Senior Vice President            2001   484,069    258,851        1,794              0    135,000          0       14,873
and President, Applied           2000   453,000    533,012        7,931              0    318,800    299,500    7,192,212
Biosystems Group

Kathy P. Ordonez(5)...........   2002   374,424    420,325            0        774,800    213,000          0       16,185
Senior Vice President            2001   196,548    167,712            0              0    218,500          0       11,916
and President, Celera
Genomics Group and
Celera Diagnostics

William B. Sawch..............   2002   405,231    298,337        2,125              0    153,000          0       81,725
Senior Vice President            2001   376,739    177,645        1,794              0    112,500          0       18,328
and General Counsel              2000   358,588    346,119        7,081              0    293,800          0    6,403,586

Dennis L. Winger..............   2002   463,846    341,996        2,125              0    153,000          0       95,114
Senior Vice President            2001   427,459    201,550        1,794      4,015,688    112,500          0       20,043
and Chief Financial              2000   410,083    392,832      136,079              0    293,800          0    7,082,372
Officer


------------------------------
  (1) Amount shown for fiscal year 2002 for Mr. White includes the incremental
      cost to the Company of providing various reportable perquisites and
      personal benefits, including the personal use of Company aircraft by
      Mr. White of $155,012, and dividend equivalents paid in cash on
      performance units. Amounts shown for fiscal 2002 for the other officers
      represent dividend equivalents paid in cash on performance units.

  (2) The dollar value of restricted stock awarded in fiscal year 2002 is based
      on the closing price of a share of Applera-Celera Genomics stock on
      April 19, 2002, the date of the grant. The grant to Ms. Ordonez consisted
      of 40,000 restricted shares of Applera-Celera Genomics stock. Of the
      shares granted, 10,000 vest on each of April 19, 2003 and April 19, 2004,
      and 20,000 vest on April 19, 2005, subject in each case to Ms. Ordonez
      being an employee of the Company as of the vesting date. As of June 30,
      2002, Mr. White held 120,000 restricted shares of Applera-Applied
      Biosystems stock and 30,000 restricted shares of Applera-Celera Genomics
      stock having an aggregate value of $2,698,800, Mr. Winger held 24,000
      restricted shares of Applera-Applied Biosystems stock and 6,000 restricted
      shares of Applera-Celera Genomics stock having an aggregate value of
      $539,760, and Ms. Ordonez held 40,000 restricted shares of Applera-Celera
      Genomics stock having an aggregate value of $480,000. As of that date,
      none of the other Named Executive Officers held any shares of restricted
      stock. Prior to vesting, Messrs. White and Winger and Ms. Ordonez have the
      right to receive dividends, if any, on and to vote, but may not sell or
      otherwise dispose of, the restricted shares.

                                       16

  (3) A breakdown of the options set forth in the table, by class, is as
      follows:



                                                            Applera-Applied           Applera-Celera
                                                            Biosystems stock          Genomics stock
                                                            ----------------          --------------
                                                                          
Mr. White......................................  2002           335,000                    85,000
                                                 2001           270,000                    67,500
                                                 2000           637,600                    75,000
Dr. Hunkapiller................................  2002           141,000                    34,000
                                                 2001           108,000                    27,000
                                                 2000           288,800                    30,000
Ms. Ordonez....................................  2002            85,000                   128,000
                                                 2001           167,200                    51,300
Mr. Sawch......................................  2002           125,000                    28,000
                                                 2001            90,000                    22,500
                                                 2000           268,800                    25,000
Mr. Winger.....................................  2002           125,000                    28,000
                                                 2001            90,000                    22,500
                                                 2000           268,800                    25,000


(4)   The amounts shown for fiscal year 2002 include (1) payment of a portion of
     the cash value of vested performance units under the Company's Performance
    Unit Bonus Plan for Mr. White, Dr. Hunkapiller, Ms. Ordonez, Mr. Sawch, and
    Mr. Winger of $125,000, $62,500, $0, $62,500, and $62,500, respectively;
    (2) the Company's contributions under the Company's Employee Savings Plan
    for Mr. White, Dr. Hunkapiller, Ms. Ordonez, Mr. Sawch, and Mr. Winger of
    $10,200, $8,204, $13,600, $10,200, and $10,200, respectively; and
    (3) amounts accrued under the savings plan component of the Company's Excess
    Benefit Plan for Mr. White, Dr. Hunkapiller, Ms. Ordonez, Mr. Sawch, and
    Mr. Winger of $20,492, $7,186, $2,585, $9,025, and $22,414, respectively.
    The amount for Dr. Hunkapiller also includes an inventor award of $915.

(5)   Ms. Ordonez became an employee of the Company on December 1, 2000.

OPTION GRANT TABLES

    The following table sets forth information regarding stock option grants of
Applera-Applied Biosystems stock to the Named Executive Officers during fiscal
year 2002.



                                     INDIVIDUAL GRANTS
-------------------------------------------------------------------------------------------     POTENTIAL REALIZABLE VALUE AT
                                 NUMBER OF       PERCENT OF TOTAL                                  ASSUMED ANNUAL RATES OF
                                 SECURITIES     OPTIONS GRANTED TO                                STOCK PRICE APPRECIATION
                                 UNDERLYING        EMPLOYEES IN      EXERCISE                        FOR OPTION TERM(2)
                                  OPTIONS             FISCAL           PRICE     EXPIRATION   ---------------------------------
NAME                           GRANTED (#)(1)       YEAR 2002         ($/SH)        DATE          5%($)              10%($)
----                           --------------   ------------------   ---------   ----------   --------------      -------------
                                                                                                
Tony L. White................         40,000(3)        0.44             25.00      8/16/11           628,895          1,593,743
                                      40,000(4)        0.44             25.00      8/16/11           628,895          1,593,743
                                     255,000           2.79             20.85       4/8/12         3,343,676          8,473,530
Michael W. Hunkapiller.......         20,000(3)        0.22             25.00      8/16/11           314,448            796,871
                                      20,000(4)        0.22             25.00      8/16/11           314,448            796,871
                                     101,000           1.10             20.85       4/8/12         1,324,358          3,356,183
Kathy P. Ordonez.............         85,000           0.93             20.85       4/8/12         1,114,559          2,824,510
William B. Sawch.............         20,000(3)        0.22             25.00      8/16/11           314,448            796,871
                                      20,000(4)        0.22             25.00      8/16/11           314,448            796,871
                                      85,000           0.93             20.85       4/8/12         1,114,559          2,824,510
Dennis L. Winger.............         20,000(3)        0.22             25.00      8/16/11           314,448            796,871
                                      20,000(4)        0.22             25.00      8/16/11           314,448            796,871
                                      85,000           0.93             20.85       4/8/12         1,114,559          2,824,510
-------------------------------------------------------------------------------------------------------------------------------
All Stockholders(5)..........                                                                   $2.7 billion       $6.8 billion
-------------------------------------------------------------------------------------------------------------------------------


------------------------------
  (1) Stock options were granted with an exercise price equal to the fair market
      value of a share of Applera-Applied Biosystems stock on the date of grant
      and, except as described below, are exercisable in four equal annual
      installments commencing on the first anniversary of the date of grant.

  (2) The values shown assume that the price of a share of Applera-Applied
      Biosystems stock will appreciate at the annual rates shown. These rates
      are arbitrarily assumed rates established by the SEC and are not intended
      as a forecast of future appreciation. The actual gain, if any, realized by
      the recipient will depend on the actual performance of the Applera-Applied
      Biosystems stock.

                                       17

  (3) Options will vest on August 16, 2004 and were granted in conjunction with
      a grant of an equal number of performance units under the Company's
      Performance Unit Bonus Plan. The performance units provide for vesting
      upon Applera-Applied Biosystems stock attaining and maintaining price
      targets of $30.00 and $35.00 or more per share for a period of 90 days.
      Performance units are payable over a minimum of eight quarters, subject to
      continued employment and an aggregate annual program expense limitation,
      in cash or Applera-Applied Biosystems stock equal to $25.00 for each
      performance unit. Prior to payment, holders receive dividend equivalents
      at the same time as, and in an amount per unit equal to, dividends paid on
      each share of Applera-Applied Biosystems Stock, but do not have any voting
      rights with respect to such performance units or the right to sell or
      otherwise dispose of such units.

  (4) Options will vest on earlier of (a) August 16, 2006 or (b) two years after
      the stock price targets referred to below have been attained. The options
      were granted in conjunction with a grant of an equal number of performance
      units under the Company's Performance Unit Bonus Plan. The performance
      units provide for vesting upon Applera-Applied Biosystems stock attaining
      and maintaining specified price targets of $40.00 and $45.00 or more per
      share for a period of 90 days. Performance units are payable in cash or
      Applera-Applied Biosystems stock equal to $25.00 for each performance
      unit. The performance units were otherwise granted on the same terms as
      those described in footnote 3 above.

  (5) These amounts represent the increase in the aggregate market value of the
      Applera-Applied Biosystems stock outstanding as of June 28, 2002, the last
      trading day of the fiscal year, assuming the annual rates of stock price
      appreciation set forth above over the ten-year period used for the Named
      Executive Officers.

    The following table sets forth information regarding stock option grants of
Applera-Celera Genomics stock to the Named Executive Officers during fiscal year
2002.



                                     INDIVIDUAL GRANTS
--------------------------------------------------------------------------------------------   POTENTIAL REALIZABLE VALUE AT
                                  NUMBER OF       PERCENT OF TOTAL                                ASSUMED ANNUAL RATES OF
                                  SECURITIES     OPTIONS GRANTED TO                               STOCK PRICE APPRECIATION
                                  UNDERLYING        EMPLOYEES IN      EXERCISE                       FOR OPTION TERM(2)
                                   OPTIONS             FISCAL           PRICE     EXPIRATION   ------------------------------
NAME                            GRANTED (#)(1)       YEAR 2002         ($/SH)        DATE         5%($)            10% ($)
----                            --------------   ------------------   ---------   ----------   ------------      ------------
                                                                                               
Tony L. White.................         85,000               3.10       18.895       4/8/12        1,010,052         2,559,670
Michael W. Hunkapiller........         34,000               1.24       18.895       4/8/12          404,021         1,023,868
Kathy P. Ordonez..............         28,000               1.02       18.895       4/8/12          332,723           843,185
                                       75,000               2.74       19.475      4/19/12          918,579         2,327,860
                                       12,500(3)            0.46       19.475      4/19/12          153,097           387,977
                                       12,500(4)            0.46       19.475      4/19/12          153,097           387,977
William B. Sawch..............         28,000               1.02       18.895       4/8/12          332,723           843,185
Dennis L. Winger..............         28,000               1.02       18.895       4/8/12          332,723           843,185
-----------------------------------------------------------------------------------------------------------------------------
All Stockholders(5)...........                                                                 $518 million      $1.3 billion
-----------------------------------------------------------------------------------------------------------------------------


------------------------------
  (1) Stock options were granted with an exercise price equal to the fair market
      value of a share of Applera-Celera Genomics stock on the date of grant
      and, except as described below, are exercisable in four equal annual
      installments commencing on the first anniversary of the date of grant.

  (2) The values shown assume that the price of a share of Applera-Celera
      Genomics stock will appreciate at the annual rates shown. These rates are
      arbitrarily assumed rates established by the SEC and are not intended as a
      forecast of future appreciation. The actual gain, if any, realized by the
      recipient will depend on the actual performance of the Applera-Celera
      Genomics stock.

  (3) Options will vest on April 19, 2005 and were granted in conjunction with a
      grant of an equal number of performance units under the Company's
      Performance Unit Bonus Plan. The performance units provide for vesting
      upon Applera-Celera Genomics stock attaining and maintaining price targets
      of $24.475 and $29.475 or more per share for a period of 90 days.
      Performance units are payable over a minimum of eight quarters, subject to
      continued employment and an aggregate annual program expense limitation,
      in cash or Applera-Celera Genomics stock equal to $19.475 for each
      performance unit. Prior to payment, holders receive dividend equivalents
      at the same time as, and in an amount per unit equal to, dividends paid on
      each share of Applera-Celera Genomics stock, if any, but do not have any
      voting rights with respect to such performance units or the right to sell
      or otherwise dispose of such units.

  (4) Options will vest on earlier of (a) April 19, 2007 or (b) two years after
      the stock price targets referred to below have been attained. The options
      were granted in conjunction with a grant of an equal number of performance
      units under the Company's Performance Unit Bonus Plan. The performance
      units provide for vesting upon Applera-Celera Genomics stock attaining and
      maintaining specified price targets of $34.475 and $39.475 or more per
      share for a period of 90 days. Performance units are payable in cash or
      Applera-Celera Genomics stock equal to $19.475 for each performance unit.
      The performance units were otherwise granted on the same terms as those
      described in footnote 3 above.

  (5) These amounts represent the increase in the aggregate market value of the
      Applera-Celera Genomics stock outstanding as of June 28, 2002, the last
      trading day of the fiscal year, assuming the annual rates of stock price
      appreciation set forth above over the ten-year period used for the Named
      Executive Officers.

                                       18

OPTION EXERCISES AND YEAR-END VALUE TABLES

    The following table sets forth information regarding the exercise of options
for Applera-Applied Biosystems stock by the Named Executive Officers during
fiscal year 2002 and the value of their unexercised options for Applera-Applied
Biosystems stock at June 30, 2002.



                                                                           NUMBER OF SECURITIES
                                                                                UNDERLYING               VALUE OF UNEXERCISED
                                                                            UNEXERCISED OPTIONS          IN-THE-MONEY OPTIONS
                                                                             AT JUNE 30, 2002              AT JUNE 30, 2002
                                                                                    (#)                         ($)(1)
                                     SHARES ACQUIRED                    ---------------------------   ---------------------------
                                       ON EXERCISE     VALUE REALIZED
NAME                                       (#)              ($)         EXERCISABLE   UNEXERCISABLE   EXERCISABLE   UNEXERCISABLE
----                                 ---------------   --------------   -----------   -------------   -----------   -------------
                                                                                                  
Tony L. White......................      189,340          3,983,415      1,731,993      1,100,100      5,447,276              0
Michael W. Hunkapiller.............        8,136            208,643        877,136        480,800      4,036,323              0
Kathy P. Ordonez...................            0                  0         31,250        220,950              0              0
William B. Sawch...................            0                  0        523,955        436,300      1,692,366              0
Dennis L. Winger...................            0                  0        631,273        436,300      1,489,284              0


------------------------------
  (1) The fair market value of a share of Applera-Applied Biosystems stock on
      June 28, 2002, the last trading day of the fiscal year, was $20.345.

    The following table sets forth information regarding the exercise of options
for Applera-Celera Genomics stock by the Named Executive Officers during fiscal
year 2002, and the value of their unexercised options for Applera-Celera
Genomics stock at June 30, 2002.



                                                                           NUMBER OF SECURITIES
                                                                                UNDERLYING               VALUE OF UNEXERCISED
                                                                            UNEXERCISED OPTIONS         IN-THE-MONEY OPTIONS AT
                                                                             AT JUNE 30, 2002                JUNE 30, 2002
                                                                                    (#)                         ($)(1)
                                     SHARES ACQUIRED                    ---------------------------   ---------------------------
                                       ON EXERCISE     VALUE REALIZED
NAME                                       (#)              ($)         EXERCISABLE   UNEXERCISABLE   EXERCISABLE   UNEXERCISABLE
----                                 ---------------   --------------   -----------   -------------   -----------   -------------
                                                                                                  
Tony L. White......................      70,496          1,581,046        687,426        321,111       2,270,089       441,775
Michael W. Hunkapiller.............       2,034             44,721        430,810        154,292       1,731,252       260,327
Kathy P. Ordonez...................           0                  0         12,825        166,475               0             0
William B. Sawch...................           0                  0        266,136        115,319         963,938       175,129
Dennis L. Winger...................           0                  0        287,231        115,319       1,002,818       175,129


------------------------------
(1)   The fair market value of a share of Applera-Celera Genomics stock on
     June 28, 2002, the last trading day of the fiscal year, was $11.795.

RETIREMENT BENEFITS

    The Company maintains a qualified defined benefit Employee Pension Plan
covering all of its domestic employees hired by the Company prior to July 1,
1999, including the Named Executive Officers other than Ms. Ordonez, and a
non-qualified Excess Benefit Plan, which provides benefits that would otherwise
be denied participants by reason of certain limitations of the Internal Revenue
Code on qualified plan benefits. The Employee Pension Plan and the Excess
Benefit Plan provide annual benefits at normal retirement age (65) based on a
participant's final average base salary (measured over 36 months from
October 1, 1995) and service from October 1, 1995 and a participant's final
average incentive compensation awards (measured over 36 months from July 1,
1995) and service from July 1, 1995. The Company also has a frozen non-qualified
Supplemental Retirement Plan, which provides benefits based on service and
awards to July 1, 1995.

    Benefits for service prior to October 1, 1995 under the Employee Pension
Plan and the pension plan component of the Excess Benefit Plan were based on a
career average benefit formula providing 1.4% of base earnings during the period
of participation, plus 0.5% of base earnings above a specified wage base called
"covered compensation" (defined by the Internal Revenue Service as a function of

                                       19

year of birth). After August 1, 1989, plan accruals for service over 35 years
were calculated at a rate of 1.7% of base earnings. A variable annuity option
was available to each participant for benefit accruals prior to October 1, 1995.
The benefit under the Supplemental Retirement Plan for service prior to July 1,
1995 was based on a career average formula providing 1.5% of the sum of all
payments made to a participant during his or her participation in the Company's
incentive compensation program.

    The Employee Pension Plan and the pension plan component of the Excess
Benefit Plan were amended in June 1999 to exclude employees hired on or after
July 1, 1999 and to cease benefit accruals after June 30, 2004.

    The following table shows the estimated total annual benefit from all the
plans payable to a covered participant at normal retirement age, for service
between October 1, 1995 and June 30, 2004.

                               PENSION PLAN TABLE



                          YEARS OF SERVICE BETWEEN OCTOBER 1, 1995 AND JUNE 30, 2004
   AVERAGE ANNUAL       --------------------------------------------------------------
    REMUNERATION            5            6            7            8           8.75
---------------------   ----------   ----------   ----------   ----------   ----------
                                                             
     $  400,000          $ 33,660     $ 40,392     $ 47,124     $ 53,856     $ 58,905
        600,000            50,780       60,936       71,092       81,248       88,865
        800,000            67,380       80,856       94,332      107,808      117,915
      1,000,000            83,500      100,200      116,900      133,600      146,125
      1,200,000           100,500      120,600      140,700      160,800      175,875
      1,400,000           117,500      141,000      164,500      188,000      205,625
      1,600,000           134,500      161,400      188,300      215,200      235,375
      1,800,000           151,500      181,800      212,100      242,400      265,125
      2,000,000           168,500      202,200      235,900      269,600      294,875
      2,200,000           185,500      222,600      259,700      296,800      324,625
      2,400,000           202,500      243,000      283,500      324,000      354,375
      2,600,000           219,500      263,400      307,300      351,200      384,125
      2,800,000           236,500      283,800      331,100      378,400      413,875
      3,000,000           253,500      304,200      354,900      405,600      443,625


    The benefit amounts shown in the Pension Plan Table are computed on a
straight life annuity basis, payable at age 65, and assume covered compensation
for social security purposes of $60,000 in all cases. As of June 30, 2002,
Mr. White, Dr. Hunkapiller, and Mr. Sawch each had 6.75 years of credited
service from October 1, 1995 for pension purposes, and Mr. Winger had
4.75 years of credited service for pension purposes. Ms. Ordonez is not a
participant in the Employee Pension Plan. The base salary and incentive
compensation for each such person are as set forth in the salary and bonus
columns of the Summary Compensation Table above under "EXECUTIVE COMPENSATION."

    Estimated annual benefits accrued prior to October 1, 1995 and payable upon
retirement at age 65 under the Employee Pension Plan, the Supplemental
Retirement Plan, and the pension plan component of the Excess Benefit Plan to
Mr. White, Dr. Hunkapiller, and Mr. Sawch are $267, $22,037, and $22,655,
respectively, assuming continued service for benefit eligibility and based on
the current variable unit value, as applicable. Mr. Winger and Ms. Ordonez do
not have benefit accruals prior to October 1, 1995.

    Under the terms of his employment agreement, Mr. White is entitled to
receive a supplemental retirement benefit equal to the annual benefit he would
have received if he were credited with 26 years of service (in addition to his
benefit for actual service) under the Employee Pension Plan and non-qualified
plans (the Excess Benefit Plan and the Supplemental Retirement Plan), reduced by
the

                                       20

annual benefit he will receive from the Employee Pension Plan and non-qualified
plans based on his actual service and further reduced by $111,528.

    The Employee Pension Plan preserves and protects the benefits of any active
participant in the plan whose employment by the Company is terminated within
three years following a change in control of the Company. In the event of such a
termination, the rights, expectancies, and the benefits of such participants (as
in effect on the date of the change in control) may not be diminished through
amendment or termination of the Employee Pension Plan after the change in
control. In addition, in the event the Employee Pension Plan is terminated
within three years following a change in control, any funds remaining after the
satisfaction of all liabilities under the plan will be allocated among
participants in accordance with applicable United States Department of Labor
regulations.

EMPLOYMENT CONTRACTS, TERMINATION OF EMPLOYMENT, CHANGE-IN-CONTROL, AND OTHER
  AGREEMENTS

    We entered into a three-year employment agreement with Mr. White dated
September 12, 1995, as amended, pursuant to which he serves as Chairman,
President and Chief Executive Officer of the Company. The agreement is
automatically extended for consecutive one-year periods unless either party
gives at least 180 days notice of its intent not to renew. No such notice has
been given by either party. Under the terms of the agreement, Mr. White receives
a base annual salary (currently set at $1,000,000) subject to annual review and
a target incentive payment (currently set at 130% of his base annual salary). In
the event of termination by the Company of Mr. White's employment without cause
or if he terminates employment for good reason (as defined), Mr. White will
receive three times his base salary and target bonus, the fair market value of
36,000 shares of Applera-Applied Biosystems stock and 18,000 shares of
Applera-Celera Genomics stock, a pro rated incentive payment, and other
specified benefits.

    We entered into an agreement with Mr. Winger dated June 3, 1997, as amended,
pursuant to which he serves as Senior Vice President and Chief Financial Officer
of the Company. Under the terms of the agreement, Mr. Winger receives a base
annual salary (currently set at $505,000) subject to annual review and a target
incentive payment (currently set at 75% of his base annual salary). He is also
eligible to receive an annual restricted stock award which vests based on
performance criteria related to the Company's cash flow. The agreement also
provides that if Mr. Winger is terminated other than for cause he will receive
two times his base salary and continuation of health benefits.

    We have agreements with each of the Named Executive Officers and certain
other key employees which provide that if, following a change in control, any of
such persons leaves employment for good reason (as defined) or his or her
employment is terminated without cause, he or she will generally be entitled to
receive between one and three times (three times in the case of each of the
Named Executive Officers) his or her base salary and incentive compensation,
full vesting of all restricted stock and stock options, and other specified
benefits.

    We have entered into deferred compensation contracts with Dr. Hunkapiller
and Mr. Sawch which, subject to certain conditions, provide for annual payments
of $25,000 to be made for a maximum of ten years, commencing on retirement from
the Company (or in the event of termination of employment for good reason (as
defined) or without cause following a change in control of the Company). The
annual payment may be reduced or forfeited if the recipient elects one of
several optional forms of payment based on actuarial determinations, terminates
employment prior to normal retirement age, or competes with the Company.

    From time to time we have provided loans to our employees, including our
executive officers, in connection with their hiring or the relocation of their
principal residence at our request. Vikram Jog, Corporate Controller, received
three such loans upon joining the Company and relocating to Connecticut. The
first of these loans, in the principal amount of $100,000, was repaid in
July 2002.

                                       21

Interest on this loan was payable annually, in arrears, at the federal short
term rate, if any, required by Section 7872 of the Internal Revenue Code, and
the principal was due on September 30, 2004. The other two loans, each in the
principal amount of $50,000, are interest-free and are forgivable in five equal
annual installments provided Mr. Jog continues to be an employee of the Company
on the installment date. If Mr. Jog voluntarily terminates his employment with
the Company or is terminated for cause, the remaining balances of the two
$50,000 loans accelerate. The largest aggregate amount outstanding on these
loans, including the repaid loan, during fiscal year 2002 was $180,000. The
aggregate amount outstanding on the two $50,000 loans at August 26, 2002 was
$60,000.

    In addition, Ugo D. DeBlasi, Assistant Controller and Vice
President--Finance of the Celera Genomics group, received two interest-free
loans in connection with his relocation from Connecticut to Maryland. The first
of these loans, in the principal amount of $100,000, is forgivable in five equal
annual installments provided Mr. DeBlasi continues to be an employee of the
Company on the installment date, and the other, in the principal amount of
$50,000, is payable on July 1, 2004. The loans are subject to acceleration if
Mr. DeBlasi ceases to be an employee of the Company. The largest aggregate
amount outstanding on these loans during fiscal year 2002 was $130,000, and the
aggregate amount outstanding on these loans at August 26, 2002 was $90,000.

    We entered into consulting arrangements with Dr. Timothy J. Hunkapiller
pursuant to which he provides services to the Applied Biosystems group in the
fields of bioinformatics and high speed text analysis. Dr. Timothy J.
Hunkapiller is the brother of Dr. Michael W. Hunkapiller, Senior Vice President
and President, Applied Biosystems group. During fiscal year 2002, we paid
$110,726 (including $27,926 in expenses) to Dr. Timothy J. Hunkapiller and an
additional $62,500 to Discovery Biosciences Corporation for these services.
Discovery Biosciences is a private corporation of which Dr. Timothy J.
Hunkapiller is president and chief scientific officer and Dr. David T. Kingsbury
was, until January 2002, chairman and chief executive officer. Dr. Kingsbury is
the husband of Deborah A. Smeltzer, a Vice President of the Company. We have
been advised that Dr. Kingsbury did not receive any cash compensation from
Discovery Biosciences during fiscal year 2002. In July 2002, we entered into a
new agreement with Dr. Timothy J. Hunkapiller pursuant to which he will assist
the Applied Biosystems group in the development of technology for the study of
biological macromolecules. Under the terms of this agreement, Dr. Timothy J.
Hunkapiller will be paid a fee of $1,400 per day (plus reasonable travel
expenses) for a minimum of 48 days during fiscal year 2003.

    In fiscal year 1999, we entered into several agreements with The Institute
for Genomic Research ("TIGR"), a Maryland non-stock corporation, in connection
with the establishment of operations at the Celera Genomics group. These
agreements included a non-exclusive license agreement for access to TIGR's Human
Gene Index database and the use of certain software developed by TIGR. We also
granted TIGR an option to purchase 2,584,700 shares of Applera-Celera Genomics
stock (unexercised as to 1,432,200 shares as of August 26, 2002) at an exercise
price of $6.42 per share in exchange for TIGR's agreement not to compete with
the Celera Genomics group in specified areas for a one year period. The option
expires on June 30, 2009. During fiscal year 2002, the Applied Biosystems group
recognized revenues of $4.7 million from the sale of instruments and consumables
to TIGR. Dr. J. Craig Venter, who resigned as Senior Vice President and
President of the Celera Genomics group in January 2002, was also Chairman of the
Board of Trustees of TIGR, and his wife, Claire M. Fraser, was President of
TIGR.

      PROPOSAL 2--RATIFICATION OF THE SELECTION OF INDEPENDENT ACCOUNTANTS

    The Audit/Finance Committee of the Board of Directors has selected
PricewaterhouseCoopers LLP ("PwC"), independent accountants, to audit the books,
records, and accounts of the Company and its subsidiaries for the fiscal year
ending June 30, 2003. We are asking you to ratify this selection at the meeting.

                                       22

    PwC has audited the Company's books annually since 1944, has offices in or
convenient to the localities in the United States and foreign countries where
the Company or its subsidiaries operate, and is considered to be well qualified.
If the selection of PwC is not ratified, the selection of independent
accountants will be reconsidered by the Audit/Finance Committee.

    The fees billed by PwC for services performed during fiscal year 2002 are
set forth below.

    AUDIT FEES.  PwC has audited the consolidated financial statements of the
Company as of and for the fiscal year ended June 30, 2002 and reviewed the
Company's quarterly reports on Form 10-Q for that fiscal year. The aggregate
fees billed by PwC for such professional services were $1,470,000. In addition,
PwC billed the Company $45,000 during fiscal 2002 for audit services performed
in the prior fiscal year.

    FINANCIAL INFORMATION SYSTEMS DESIGN AND IMPLEMENTATION FEES.  PwC did not
provide the Company with any professional services in connection with Financial
Information Systems Design and Implementation during fiscal 2002.

    ALL OTHER FEES.  The aggregate fees billed by PwC for services rendered to
the Company, other than the services described above under "Audit Fees" and
"Financial Information Systems Design and Implementation Fees," during fiscal
year 2002 were $1,829,500. These services included tax compliance and advisory
services and assistance with acquisition activities and other matters. In
addition, PwC billed the Company $28,000 during fiscal year 2002 for such
services performed in the prior fiscal year.

    A representative of PwC will attend the meeting, have the opportunity to
make a statement, and be available to respond to appropriate questions.

    THE BOARD RECOMMENDS THAT YOU VOTE "FOR" THIS PROPOSAL.

  PROPOSAL 3--APPROVAL OF AMENDMENTS TO THE APPLERA CORPORATION 1999 EMPLOYEE
                              STOCK PURCHASE PLAN

    Proposal 3 pertains to amendments to the Applera Corporation 1999 Employee
Stock Purchase Plan (the "Purchase Plan"). The sole purpose of these amendments
is to increase the number of shares of Applera common stock authorized for
issuance under the Purchase Plan. We are asking you to approve these amendments
at the meeting.

    For more than 40 years the Company has maintained a series of employee stock
purchase plans to provide employees with an incentive to acquire an equity
interest in the Company and to work effectively for its growth and prosperity. A
substantial number of the Company's employees have become, and continue to be,
stockholders of the Company as a result of these plans. The Board believes that
these plans have generally been beneficial to employees as well as the Company
and its stockholders because they have enhanced the interest of the employees in
the continued success of the Company and further aligned the interests of
employees and stockholders. In addition, the Board is of the opinion that
employee stock purchase plans provide an effective aid in recruiting highly
qualified and talented employees.

    The Purchase Plan was approved by the stockholders at the 1999 Annual
Meeting and provides for offerings during each of the five calendar years
commencing January 1, 2000. As of June 30, 2002, there remained available for
issuance under the Purchase Plan approximately 1,290,000 shares of
Applera-Applied Biosystems stock out of a total of 2,000,000 authorized and
approximately 460,000 shares of Applera-Celera Genomics stock out of a total of
1,200,000 authorized. As a result of the level of past employee participation in
the Purchase Plan, the Board believes that more shares of Applera-

                                       23

Applied Biosystems stock and Applera-Celera Genomics stock will be necessary to
continue to offer such shares through the term of the Purchase Plan. If the
amendments are not approved by the stockholders, they will not be effective.

SUMMARY OF PROPOSED AMENDMENTS

    The proposed amendments would increase the number of shares of
Applera-Applied Biosystems stock authorized for issuance under the Purchase Plan
by 1,000,000 and increase the number of shares of Applera-Celera Genomics stock
authorized for issuance under the Purchase Plan by 1,500,000. We are not seeking
an extension of the term of the Purchase Plan.

SUMMARY OF THE PURCHASE PLAN

    This summary highlights all material information from the Purchase Plan as
proposed to be amended. If you would like to obtain a complete copy of the
Purchase Plan, please contact the Corporate Secretary at 301 Merritt 7,
P.O. Box 5435, Norwalk, CT 06856-5435, telephone (203) 840-2000.

    SHARES SUBJECT TO THE PLAN; TERM.  Subject to adjustment as provided below,
if the amendments to the Purchase Plan are approved, an additional 1,000,000
shares of Applera-Applied Biosystems stock and an additional 1,500,000 shares of
Applera-Celera Genomics stock will be available for issuance under the Purchase
Plan. Shares delivered under the Purchase Plan may be newly issued shares or
treasury shares. See "EQUITY COMPENSATION PLAN INFORMATION" under Proposals 4
and 5 below for more information about Applera common stock authorized for
issuance under our equity compensation plans.

    No offering may be made under the Purchase Plan after December 31, 2004. As
of August 26, 2002, the fair market value of a share of Applera-Applied
Biosystems stock was $20.625 and the fair market value of a share of
Applera-Celera Genomics stock was $10.45.

    ADMINISTRATION.  The Purchase Plan is administered by the Management
Resources Committee of the Board (the "Committee"). The Committee has the
discretion to interpret the Purchase Plan and make rules and regulations
relating thereto. None of the members of the Committee are eligible to
participate in the Purchase Plan.

    PARTICIPATION.  All employees of the Company and designated subsidiaries on
the date of each offering are eligible to participate in the Purchase Plan.
Directors who are not employees of the Company and any person who, after the
grant of an option to purchase, would hold 5% or more of the total combined
voting power or value of the Company are not eligible to participate. As of
July 31, 2002, approximately 4,000 United States and 1,700 non-United States
employees of the Company were eligible to participate in the Purchase Plan. Up
to an additional approximately 150 non-United States employees would be eligible
to participate in the Purchase Plan if the Committee were to designate all
majority-owned subsidiaries of the Company as participating subsidiaries.

    PURCHASES UNDER THE PURCHASE PLAN.  The Company makes quarterly offerings to
eligible employees in the United States to purchase shares of Applera-Applied
Biosystems stock and Applera-Celera Genomics stock. The offerings relate to four
three-month purchase periods commencing on the first trading day of January,
April, July, and October of each year during the term of the Purchase Plan. Each
eligible employee is offered the right to purchase shares of Applera-Applied
Biosystems stock and/or Applera-Celera Genomics stock at a purchase price for
each class of stock equal to the lower of 85% of the fair market value of the
class of stock on the first day of the purchase period or 85% of the fair market
value of the class of stock on the last day of the purchase period.

                                       24

    Purchases under the Purchase Plan are made automatically at the end of each
purchase period using payroll deductions during the purchase period authorized
by the participating employees. Participating employees can set aside up to 10%
of their compensation in each purchase period for the purchase of
Applera-Applied Biosystems stock and/or Applera-Celera Genomics stock. No
eligible employee can elect a payroll deduction at a rate that would cause him
or her to purchase more than $25,000 of Applera common stock in any calendar
year.

    Offerings to non-United States employees are made no more frequently than
annually and the terms of such offerings may otherwise vary depending on local
custom and law.

    A participating employee has none of the rights or privileges of a
stockholder of the Company (including the right to receive dividends) until the
shares purchased under the Purchase Plan have been fully paid for and issued.

    WITHDRAWAL.  A participating employee may during any purchase period
(1) direct the Company to make no further deductions from his or her
compensation or (2) cancel his or her option to purchase shares of stock at the
end of that purchase period. If the employee has directed that payroll
deductions be discontinued, any sums deducted will be retained by the Company
until the end of the purchase period, at which time the employee will receive
that number of whole and fractional shares which can be purchased with the
amount so retained. If the employee has directed that the option be canceled,
the Company will, as soon as practicable thereafter, refund in cash, without
interest, all amounts credited to the employee's account.

    TERMINATION OF EMPLOYMENT.  If the employment of a participating employee is
terminated prior to the end of a purchase period because of total and permanent
disability, retirement, or death, the employee or his or her legal
representative may either (1) cancel the option to purchase and, as soon as
practicable thereafter, receive a cash refund, without interest, of all payroll
deductions credited to his or her account with respect to the offering, or
(2) elect to receive at the end of the purchase period that number of whole and
fractional shares of stock which such payroll deductions will purchase.

    If the employment of a participating employee is terminated during a
purchase period for any other reason, he or she will receive, as soon as
practicable thereafter, a cash refund, without interest, of all payroll
deductions credited to his or her account with respect to the offering.

    TRANSFERABILITY.  A participating employee's rights under the Purchase Plan
are exercisable, during his or her lifetime, only by the employee and may not be
transferred in any manner. After the death of a participating employee, his or
her rights may be transferred pursuant to the laws of descent and distribution.

    AWARDS IN FOREIGN COUNTRIES.  The Committee has the authority and discretion
to adopt procedures and modify the Purchase Plan as it deems necessary or
desirable to comply with the provisions of the laws of foreign countries in the
which the Company operates in order to assure the viability of the benefits of
the Purchase Plan to the individuals employed in such countries and to meet the
objectives of the Purchase Plan.

    TERMINATION AND AMENDMENT.  The Committee may terminate the Purchase Plan at
any time or make any amendment or modification it deems advisable.

    ADJUSTMENTS.  The Purchase Plan provides that the Committee may adjust, as
it deems appropriate, the number and class of shares available under the
Purchase Plan to reflect changes in the outstanding stock that occur because of
stock dividends, stock splits, recapitalizations, reorganizations, liquidations,
or other similar events.

                                       25

CERTAIN FEDERAL INCOME TAX CONSEQUENCES

    The Company believes that, based on the laws as in effect on the date of
this proxy statement, the following are the principal federal income tax
consequences to participants and the Company of participation in the Purchase
Plan. THIS SUMMARY IS NOT A COMPLETE ANALYSIS OF ALL POTENTIAL TAX CONSEQUENCES
RELEVANT TO PARTICIPANTS AND THE COMPANY AND DOES NOT DESCRIBE TAX CONSEQUENCES
BASED ON PARTICULAR CIRCUMSTANCES. STATE, LOCAL, AND FOREIGN TAX LAWS ARE NOT
DISCUSSED.

    It is intended that the option to purchase shares of stock granted under the
Purchase Plan will constitute an option issued pursuant to an "employee stock
purchase plan" within the meaning of Section 423 of the Internal Revenue Code.
If shares are purchased under the Purchase Plan, no income will be realized by
the employee at the time of the transfer of the shares to such employee if (1)
the Purchase Plan constitutes an "employee stock purchase plan," (2) no
disposition of these shares is made within two years after the date of grant of
the option or within one year after the purchase of the shares, and (3) the
employee is an employee at all times from the date of grant of the option to the
date three months before exercise of the option. If an employee dies while
owning the shares, no income will be realized at the time of the transfer.
Further, when an employee or his or her estate sells or otherwise disposes of
the shares, there will be included in his, her, or its income, as compensation,
an amount equal to the lesser of:

    - the amount by which the fair market value of the shares on the first day
      of the offering period exceeds the purchase price for the shares, or

    - the amount by which the fair market value at the time of disposition or
      death exceeds the purchase price per share.

Any further gain will be treated for tax purposes as long-term capital gain,
provided that the employee holds the shares for the applicable long-term capital
gain holding period after the last day of the offering period applicable to such
shares.

    If an employee disposes of the shares within either the two-year or one-year
period referred to above, he or she will realize ordinary income in the year of
disposition in an amount equal to the difference between the purchase price and
the fair market value of the shares at the time of exercise of the option. Any
difference between the amount received upon such a disposition and the fair
market value of the shares at the time of exercise of the option will be capital
gain or loss, as the case may be.

    No deduction will be allowed to the Company for federal income tax purposes
in connection with the grant or exercise of the option to purchase shares under
the Purchase Plan, provided there is no disposition of shares by a participating
employee within either the two-year or the one-year period referred to above. If
there is a disposition of shares within either of these periods, the Company
will be entitled to a deduction in the same amount and at the same time that the
employee realizes ordinary income.

NEW PLAN BENEFITS

    Participation in the Purchase Plan is voluntary and each eligible employee
will make his or her own election whether and to what extent to participate in
the plan. It is therefore not possible to determine the benefits or amounts that
will be received in the future by individual employees or groups of employees
under the Purchase Plan.

                                       26

VOTE REQUIRED FOR APPROVAL

    Approval of this proposal requires the favorable vote of a majority of votes
present in person or by proxy and entitled to vote at the meeting.

    THE BOARD RECOMMENDS THAT YOU VOTE "FOR" THIS PROPOSAL.

                PROPOSALS 4 AND 5--APPROVAL OF AMENDMENTS TO THE
 APPLERA CORPORATION/APPLIED BIOSYSTEMS GROUP 1999 STOCK INCENTIVE PLAN AND THE
      APPLERA CORPORATION/CELERA GENOMICS GROUP 1999 STOCK INCENTIVE PLAN

    Proposal 4 pertains to an amendment to the Applera Corporation/Applied
Biosystems Group 1999 Stock Incentive Plan (the "Applera-Applied Biosystems
Plan") and Proposal 5 pertains to an amendment to the Applera Corporation/Celera
Genomics Group 1999 Stock Incentive Plan (the "Applera-Celera Genomics Plan").
These plans and amendments are identical except as noted below. (The
Applera-Applied Biosystems Plan and the Applera-Celera Genomics Plan are
sometimes referred to collectively as the "Incentive Plans.") We are asking you
to approve these amendments at the meeting.

    The sole purpose of the amendments is to increase the number of shares
authorized for issuance under each of the Incentive Plans. The Board of
Directors believes that stock-based incentives are in the best interest of the
Company and its stockholders as they tie employee interests directly to
stockholder interests. The Board also believes that stock-based incentives are
an important component of the compensation programs of life science and other
technology companies with which the Company must compete for employees. As a
result, the Board has found that stock-based incentives are necessary to attract
and retain employees throughout the Company, and stock-based incentives continue
to be a significant component of the Company's overall compensation program.
During fiscal 2002, approximately 4,600 employees, or more than 75% of the
Company's worldwide workforce, received stock options under the Incentive Plans.

    As of June 30, 2002, approximately 9,100,000 shares of Applera-Applied
Biosystems stock remained available for the grant of new awards under the
Applera-Applied Biosystems Plan out of a total of 37,600,000 authorized to be
issued under that plan, and approximately 4,000,000 shares of Applera-Celera
Genomics stock remained available for the grant of new awards under the Applera-
Celera Genomics Plan out of a total of 17,200,000 shares authorized to be issued
under that plan.

    The Board has not repriced any outstanding stock options, and as of
August 26, 2002, 81% of the outstanding stock options for shares of
Applera-Applied Biosystems stock and 61% of the outstanding stock options for
shares of Applera-Celera Genomics stock were out-of-the-money. Accordingly, the
Board believes that the new shares being requested will be particularly
important in order to provide the incentives necessary to retain and motivate
existing employees.

    If the amendments are not approved by the stockholders, awards would
continue to be made under the Incentive Plans until March 31, 2004. However, the
Board believes that it will not have sufficient shares to provide appropriate
employee incentives and to satisfy ongoing needs and anticipated growth.

SUMMARY OF PROPOSED AMENDMENTS

    The proposed amendments would increase the number of shares authorized for
issuance under the Applera-Applied Biosystems Plan by 4,000,000 and increase the
number of shares authorized for issuance under the Applera-Celera Genomics Plan
by 1,500,000. We are not seeking an extension of the term of either Incentive
Plan.

                                       27

SUMMARY OF THE INCENTIVE PLANS

    This summary highlights all material information from the Incentive Plans as
proposed to be amended. If you would like to obtain a complete copy of the
Incentive Plans, please contact the Corporate Secretary at 301 Merritt 7,
P.O. Box 5435, Norwalk, CT 06856-5435, telephone (203) 840-2000.

    SHARES SUBJECT TO THE PLANS.  Subject to adjustment as discussed below, if
the amendments are approved, an additional 4,000,000 shares of Applera-Applied
Biosystems stock will be available for issuance under the Applera-Applied
Biosystems Plan and an additional 1,500,000 shares of Applera-Celera Genomics
stock will be available for issuance under the Applera-Celera Genomics Plan. As
of August 26, 2002, the fair market value of a share of Applera-Applied
Biosystems stock was $20.625 and the fair market value of a share of
Applera-Celera Genomics stock was $10.45.

    TYPES OF INCENTIVES.  Incentives granted under the Incentive Plans may be:

    - stock options, consisting of incentive stock options within the meaning of
      Section 422 of the Internal Revenue Code and non-qualified stock options
      (collectively, "Options");

    - shares of Applera-Applied Biosystems stock or Applera-Celera Genomics
      stock, which may be subject to restrictions ("Employee Stock Awards");

    - shares of Applera-Applied Biosystems stock or Applera-Celera Genomics
      stock subject to performance goals ("Performance Shares"); or

    - director stock awards, which are shares of Applera-Applied Biosystems
      stock or Applera-Celera Genomics stock subject to restrictions ("Director
      Stock Awards").

    ELIGIBILITY.  Under the terms of the Incentive Plans:

    - all regular salaried employees, including executive officers, may receive
      Options, Employee Stock Awards, and Performance Shares;

    - all consultants performing significant services may receive non-qualified
      stock options; and

    - all non-employee directors may receive non-qualified stock options and
      Director Stock Awards.

    As of July 30, 2002, approximately 5,900 employees, consultants, and
directors were eligible to participate in the Incentive Plans.

    ADMINISTRATION.  The Incentive Plans are administered by the Management
Resources Committee (the "Committee") of the Board. The Committee determines,
subject to the terms of the Incentive Plans, the employees, non-employee
directors, and consultants to whom, and the time or times at which, it will
grant awards, as well as the terms and provisions of each award.

    STOCK OPTIONS.  The purchase price, vesting period, and all other terms and
conditions of each Option are determined by the Committee, except that the
purchase price of a share of Applera-Applied Biosystems stock or Applera-Celera
Genomics stock covered by an Option may not be less than 100% of the fair market
value of the underlying stock on the date of the grant, and the term of each
Option may not be more than ten years from the date of grant.

    If the employment of an employee, the service of a non-employee director, or
the service of a consultant to whom an Option has been granted is terminated,
other than by reason of retirement, disability, or death, the employee,
non-employee director, or consultant may exercise the Option, to the extent that
he or she is entitled to do so at the date of termination, for 30 days after the
termination, but not after the Option expires. In addition, Options held by an
employee, non-employee director, or consultant whose employment or service with
the Company is terminated for cause will be forfeited.

                                       28

"Cause" is defined as (1) any act which is in bad faith and to the detriment of
the Company or (2) a material breach of any agreement with or material
obligation to the Company.

    If an employee to whom an Option has been granted retires from the Company
under any pension plan provided by the Company or if an employee or a consultant
to whom an Option has been granted becomes totally and permanently disabled, the
Option may be fully exercised without regard to the period of continuous
employment or service at any time: (1) in the case of an incentive stock option,
within three months after retirement or disability, but not after the Option
expires; or (2) in the case of a non-qualified stock option, within one year
after retirement or disability, but not after the Option expires. If a
non-employee director (1) retires from the Board on reaching normal retirement
age, (2) resigns or declines to stand for reelection with the approval of the
Board, or (3) becomes totally and permanently disabled, the Option may be fully
exercised, without regard to the period of continuous service, at any time
within three years after retirement, resignation, declining, or disability, but
not after the Option expires.

    If an employee, non-employee director, or consultant to whom an Option has
been granted dies while employed by or engaged to provide services or while
serving as a member of the Board, the Option may be exercised to the extent that
he or she was entitled to do so at the date of death by his or her executor or
administrator or other person at the time entitled by law to the employee's,
non-employee director's, or consultant's rights under the Option. The person
exercising the Option must generally do so within one year after the death.

    Options will be exercisable only by the optionee or his or her guardian or
legal representative, and may not be transferred, except under a domestic
relations order. However, the Committee may, in its sole discretion, permit an
optionee to transfer a non-qualified stock option to certain family related
entities. After the death of an optionee, the Option may be transferred pursuant
to the laws of descent and distribution.

    A condition to the exercise of an Option following termination of employment
or service is that the optionee has not (1) rendered services or engaged
directly or indirectly in any business which, in the opinion of the Committee,
competes with or is in conflict with the interests of the Company, or
(2) violated any written agreement with the Company. An optionee's violation of
either of these conditions will result in the forfeiture of all Options held.

    Except as discussed below, no one individual may be granted an Option or
Options under either Incentive Plan during any fiscal year for an aggregate
number of shares of stock which exceeds 10% of the total number of shares
reserved for issuance under the respective Incentive Plan.

    EMPLOYEE STOCK AWARDS.  Employee Stock Awards may be subject to
restrictions, as determined by the Committee. Until those conditions are met,
the recipient may not sell, transfer, or otherwise dispose of the shares.
Recipients of Employee Stock Awards are otherwise entitled to the rights of a
stockholder with respect to the shares of stock subject to Employee Stock Awards
as the Committee may determine, including the right to vote and receive
dividends and other distributions made with respect to the stock.

    If a recipient of an Employee Stock Award terminates employment before any
applicable restrictions lapse, by reason of death, total and permanent
disability, retirement, or resignation or discharge from employment other than
for cause, the Committee may, in its sole discretion, remove restrictions on all
or a portion of the stock subject to the Employee Stock Award.

    Subject to adjustment as discussed below, no employee may receive an
Employee Stock Award under the Applera-Applied Biosystems Plan representing more
than 160,000 shares of Applera-Applied Biosystems stock during any fiscal year,
and the maximum number of shares that may be issued to all employees as Employee
Stock Awards under the Applera-Applied Biosystems Plan is 320,000. Subject to
adjustment as discussed below, no employee may receive an Employee Stock Award
under the

                                       29

Applera-Celera Genomics Plan representing more than 80,000 shares of
Applera-Celera Genomics stock during any fiscal year, and the maximum number of
shares that may be issued to all employees as Employee Stock Awards under the
Applera-Celera Genomics Plan is 160,000.

    PERFORMANCE SHARES.  Performance Shares will be subject to the attainment of
performance goals within the meaning of Section 162(m) of the Internal Revenue
Code and the regulations thereunder. These performance goals could relate to
stock price, market share, sales, earnings per share, return on equity, costs,
and cash flow, as determined by the Committee, and the period in which these
goals are to be met will not be less than one year. Certificates representing
Performance Shares will be registered in the name of the award recipient but
remain in the physical custody of the Company until the Committee has determined
that the performance goals have been attained and other stock restrictions, if
any, have been satisfied. Until Performance Shares are delivered to an award
recipient, the recipient may not sell, transfer, or otherwise dispose of those
shares. Recipients of Performance Shares are entitled to such other rights of a
stockholder with respect to Performance Shares as the Committee determines,
including the right to vote and receive dividends and other distributions.

    If a recipient of Performance Shares terminates employment by reason of
death, total and permanent disability, retirement, resignation, or discharge
from employment other than for cause before all applicable performance goals
have been attained, the Committee may, in its sole discretion, remove
restrictions on all or a portion of the Performance Shares or determine that the
performance objectives with respect to all or a portion of the Performance
Shares have been attained. However, the Committee may not exercise its
discretion to the extent that it would cause the award of Performance Shares not
to qualify as performance-based compensation under Section 162(m) of the
Internal Revenue Code.

    Subject to adjustment as discussed below, no employee may receive
Performance Shares under the Applera-Applied Biosystems Plan representing more
than 400,000 shares of Applera-Applied Biosystems stock during any fiscal year,
and the maximum number of shares that may be issued to all employees as
Performance Shares under the Applera-Applied Biosystems Plan is 1,600,000.
Subject to adjustment as discussed below, no employee may receive Performance
Shares under the Applera-Celera Genomics Plan representing more than 200,000
shares of Applera-Celera Genomics stock during any fiscal year, and the maximum
number of shares that may be issued to all employees as Performance Shares under
the Applera-Celera Genomics Plan is 800,000.

    DIRECTOR STOCK AWARDS.  As of the date of each election or reelection to the
Board, each non-employee director will automatically be granted a Director Stock
Award of up to 1,200 shares of Applera-Applied Biosystems stock under the
Applera-Applied Biosystems Plan and up to 300 shares of Applera-Celera Genomics
stock under the Applera-Celera Genomics Plan, in each case subject to adjustment
as provided below. Non-employee directors elected other than at an annual
meeting are granted a pro rata portion of such shares. Each Director Stock Award
vests on the date immediately preceding the first annual meeting of stockholders
next following the date of grant, provided that the holder continues to serve as
a member of the Board as of that date.

    The aggregate dollar value of all Director Stock Awards to a director under
both Incentive Plans could not exceed $148,500 for the fiscal year ending
June 30, 2002, and may not exceed $163,350 for the fiscal year ending June 30,
2003, increasing by 10% each year thereafter. The aggregate dollar value is
calculated by multiplying the total number of shares subject to Director Stock
Awards under both Incentive Plans by the fair market value of the applicable
class of stock on the date of grant. To the extent that the aggregate dollar
value of such awards would otherwise exceed this dollar limit, the Committee
will allocate shares of each class of stock based on the ratio of the number of
shares of each class of stock outstanding on the date of grant.

    Except as set forth below, the holder of a Director Stock Award will be
entitled to all rights of a stockholder with respect to the shares of
Applera-Applied Biosystems stock or Applera-Celera Genomics stock issued under
the Director Stock Award, including the right to receive dividends and to

                                       30

vote the shares. However, stock dividends paid on the shares will be restricted
to the same extent as the shares underlying the Director Stock Award. Prior to
vesting, the shares of stock issued under a Director Stock Award may not be
sold, transferred, or otherwise disposed of.

    If a non-employee director to whom a Director Stock Award has been granted
ceases to serve as a director as a result of death, retiring from the Board upon
reaching normal retirement age, becoming totally and permanently disabled, or
resigning with the approval of the Board, all shares subject to the Director
Stock Award will be fully vested as of the date of termination of service.

    Non-employee directors are permitted to defer receipt of their Director
Stock Awards. Deferred awards are credited to a bookkeeping account and those
awards are deemed invested in stock units, each unit representing one share of
Applera-Applied Biosystems stock or Applera-Celera Genomics stock. As dividends
are paid, a corresponding number of additional units are credited to the
director's deferral account. A non-employee director who defers receipt of a
Director Stock Award will not have voting rights with respect to the Director
Stock Award until such time as he or she receives an actual distribution of the
stock.

    CHANGE OF CONTROL.  All outstanding Options granted under the Incentive
Plans will become fully and immediately exercisable, all restrictions on
Employee Stock Awards and awards of Performance Shares will immediately
terminate, all performance objectives applicable to awards of Performance Shares
will be deemed attained, and all Director Stock Awards will become fully vested
if:

    - a tender offer or exchange offer, other than an offer by the Company, is
      made for common stock representing more than 25% of the combined voting
      power of the outstanding voting securities of the Company entitled to vote
      generally in the election of directors;

    - any person acquires common stock representing more than 25% of such
      combined voting power;

    - a majority of the incumbent directors ceases to remain on the Board; or

    - the stockholders approve the sale of all or substantially all of the stock
      or assets of the Company.

    TERMINATION AND AMENDMENT; NO REPRICING.  No award may be made under the
Incentive Plans after March 31, 2004. The Board may at any time prior to that
date terminate either of the Incentive Plans or make any amendment or
modification it deems advisable. However, any such amendments will require
stockholder approval if they would:

    - increase the aggregate number of shares which may be issued;

    - materially modify the eligibility requirements for participation; or

    - materially increase the benefits accruing to participants under either
      Incentive Plan.

    The Committee may amend the terms of any outstanding Option or other award
under either Incentive Plan at any time in its discretion in any manner it deems
appropriate, including accelerating the date of exercise of any award,
terminating restrictions, or converting an incentive stock option into a
non-qualified stock option. However, no amendment may adversely affect in any
material manner any right of any recipient without his or her consent. In
addition, the Committee may not (1) amend any previously issued award of
Performance Shares to the extent that the amendment would cause the award not to
qualify as performance-based compensation under Section 162(m) of the Internal
Revenue Code, or (2) amend any previously issued Option to reduce the purchase
price, whether by modification of the Option or by cancellation of the Option in
consideration of the immediate issuance of a replacement Option with a reduced
purchase price.

                                       31

    AWARDS IN FOREIGN COUNTRIES.  The Committee has the authority and discretion
to adopt procedures and modify the Incentive Plans as it deems necessary or
desirable to comply with the provisions of the laws of foreign countries in the
which the Company operates in order to assure the viability of the benefits of
the Options and other awards made to individuals employed in such countries and
to meet the objectives of the Incentive Plans.

    ADJUSTMENTS BY THE COMMITTEE.  The Incentive Plans provide that the
Committee may adjust, as it deems appropriate, the maximum number of shares that
may be subject to Options or other awards, and the terms of any outstanding
Options or other awards under the Incentive Plans, to reflect changes in the
outstanding stock that occur because of stock dividends, stock splits,
recapitalizations, reorganizations, liquidations, or other similar events.

CERTAIN FEDERAL INCOME TAX CONSEQUENCES

    The Company believes that, based on the laws as in effect on the date of
this proxy statement, the following are the principal federal income tax
consequences to participants and the Company of Options and other awards granted
under the Incentive Plans. THIS SUMMARY IS NOT A COMPLETE ANALYSIS OF ALL
POTENTIAL TAX CONSEQUENCES RELEVANT TO PARTICIPANTS AND THE COMPANY AND DOES NOT
DESCRIBE TAX CONSEQUENCES BASED ON PARTICULAR CIRCUMSTANCES. STATE, LOCAL, AND
FOREIGN TAX LAWS ARE NOT DISCUSSED.

    INCENTIVE STOCK OPTIONS.  If the Company issues shares to an employee upon
the exercise of an incentive stock option granted under the Incentive Plans
during the employee's employment or within three months after the employee's
termination of employment, then:

    - the employee will not recognize income at the time of the grant of the
      incentive stock option or upon exercise of the incentive stock option;

    - the Company will not be allowed a federal income tax deduction in
      connection with the grant or exercise of the incentive stock option; and

    - upon a sale or exchange of the shares after the later of (1) one year from
      the date of transfer of the shares to the employee or (2) two years from
      the date of grant of the incentive stock option, any amount received by
      the employee in excess of the incentive stock option price will be taxed
      to the employee as a capital gain, and any loss sustained by the employee
      will be a capital loss. The capital gain, if any, from sales or exchanges
      of shares is subject to tax at various rates depending upon the length of
      time the shares were held, the date of disposition, and the income tax
      bracket of the employee.

    If the shares are disposed of before the holding period requirements are
satisfied, then:

    - the employee will recognize ordinary income in the year of disposition in
      an amount (1) equal to the excess, on the date of exercise of the
      incentive stock option, of the fair market value of the shares received
      over the option price paid, but (2) limited to the excess of the amount
      received on the sale over the option price if the amount received is less
      than the fair market value on the date of exercise;

    - the Company will be entitled to a deduction for the year equal to the
      ordinary income recognized by the employee; and

    - the employee will have capital gain or loss equal to the difference
      between (1) the amount received by the employee upon the sale or exchange
      of the shares and (2) the option price paid by the employee increased by
      any ordinary income recognized.

                                       32

    NON-QUALIFIED STOCK OPTIONS.  An employee, consultant, or director to whom a
non-qualified stock option is granted will not recognize income at the time the
option is granted. When the employee, consultant, or director exercises the
option, he or she will recognize ordinary income equal to the excess, if any, of
the market value, as of the date of exercise, of the shares received over the
option price paid. Subject to the Internal Revenue Code and regulations
thereunder, the Company will generally be entitled to a federal income tax
deduction equal to the ordinary income recognized by the employee, consultant,
or director. Any compensation included in an employee's gross income will be
subject to federal and applicable state employment taxes. Upon the sale of
shares acquired through the exercise of a non-qualified stock option, the
employee, consultant, or director will have capital gain or loss equal to the
difference between (1) the amount received by the employee, consultant, or
director upon the sale or exchange of the shares and (2) the option price paid
by the employee, consultant, or director increased by any ordinary income
recognized.

    EMPLOYEE STOCK AWARDS.  No taxable income will be recognized by an employee
upon the grant of an Employee Stock Award that is subject to a substantial risk
of forfeiture unless the employee makes the election under Section 83(b) of the
Internal Revenue Code referred to in the next paragraph. If the employee does
not make an election, he or she will recognize ordinary income at the time his
or her interest in the shares is either transferable or no longer subject to a
substantial risk of forfeiture (the "Section 83 Restrictions"). The amount of
this ordinary income will be equal to the excess, if any, of the fair market
value of the shares received at the time of the lapse of the Section 83
Restrictions over the amount, if any, the employee paid for the shares. The
employee's tax basis in the shares received at the lapse of the Section 83
Restrictions will be equal to the amount, if any, paid for the shares plus the
amount of ordinary income recognized. Dividends paid on shares while they are
subject to the Section 83 Restrictions will be taxable as ordinary compensation
income and not as dividends.

    An employee receiving shares under an Employee Stock Award may elect under
Section 83(b) of the Internal Revenue Code to be taxed at the time the employee
receives the shares on an amount equal to the fair market value of the shares
received, determined without regard to the Section 83 Restrictions, at the time
of transfer less the purchase price, if any, paid for the shares. The employee's
tax basis in the shares will be the fair market value of the shares at the time
the shares are received. If a Section 83(b) election is made, dividends paid on
these shares will not be taxable as compensation income but will be taxable as
dividends, and no additional compensation income will be recognized when the
Section 83 Restrictions lapse or are released. Any compensation included in an
employee's gross income will be subject to federal and applicable state
employment taxes.

    PERFORMANCE SHARES.  No taxable income will be recognized by an employee
upon the grant of Performance Shares that is subject to a substantial risk of
forfeiture unless the employee makes the election under Section 83(b) referred
to above. If the employee does not make an election, he or she will recognize
ordinary income at the time the Performance Shares vest or are no longer subject
to a substantial risk of forfeiture. The income will be equal to the excess, if
any, of the fair market value of the shares at the time they become vested or
non-forfeitable over the amount, if any, the employee paid for the shares. If
the employee is entitled to receive dividends on the shares prior to the time
they vest or become non-forfeitable, the dividends will be taxable as ordinary
compensation income and not as dividends. The employee's tax basis in the shares
will be equal to the amount, if any, paid for the shares plus the amount of
ordinary income recognized with respect to the shares.

    An employee receiving Performance Shares may elect under Section 83(b) of
the Internal Revenue Code to be taxed at the time the employee receives the
shares on an amount equal to the fair market value of the shares received,
determined without regard to the Section 83 Restrictions, at the time of
transfer less the purchase price, if any, paid for the shares. The employee's
tax basis in the shares will be the fair market value of the shares at the time
the shares are received. If a Section 83(b) election is made, dividends paid on
the shares will not be taxable as compensation income but will be taxable as

                                       33

dividends and no additional compensation income will be recognized when the
shares vest or become non-forfeitable. Any compensation included in an
employee's gross income will be subject to federal and applicable state
employment taxes.

    DIRECTOR STOCK AWARDS.  No taxable income will be recognized by a
non-employee director upon the grant of a Director Stock Award unless he or she
makes the election under Section 83(b) referred to above. If no election is
made, the director will recognize ordinary income at the time his or her
interest in the shares vests or is no longer subject to a substantial risk of
forfeiture. The amount of ordinary income will be equal to the excess, if any,
of the fair market value of the shares received at such time over the amount, if
any, the director paid for the shares. Dividends paid on shares while they are
subject to a substantial risk of forfeiture will be taxable as ordinary income
and not as dividends. The director's tax basis in the shares received will be
equal to the amount, if any, paid for the shares plus the amount of ordinary
income recognized.

    If a Section 83(b) election is made, the director will be taxed at the time
the director receives the shares on an amount equal to the fair market value of
the shares received, determined without regard to the Section 83 Restrictions,
at the time of transfer less the purchase price, if any, paid for the shares.
The director's tax basis in the shares will be the fair market value of the
shares at the time the shares are received. If a Section 83(b) election is made,
dividends paid on the shares will not be taxable as ordinary income but will be
taxable as dividends and no additional ordinary income will be recognized when
the shares vest.

    DEFERRALS.  In general, a non-employee director who elects to defer a
Director Stock Award will not be subject to current federal income tax on the
award, or related earnings, until it is distributed. Deferred compensation
distributed under the Incentive Plans will be taxed as ordinary income and not
as capital gains.

    LIMITS ON DEDUCTIONS.  Under Section 162(m) of the Internal Revenue Code,
compensation paid to the Company's chief executive officer and the four other
most highly paid executive officers in a particular year is limited to
$1 million per person, except that compensation that is performance-based will
be excluded for purposes of calculating the amount of compensation subject to
this $1 million limitation. The Company's ability to deduct compensation paid to
any other executive officer or employee is not affected by this provision. As
noted above under "EXECUTIVE COMPENSATION--REPORT OF THE MANAGEMENT RESOURCES
COMMITTEE," the Committee generally seeks to maximize the deductibility of
compensation paid to its executive officers. However, it will maintain
flexibility to take actions that may be based on considerations other than tax
deductibility.

NEW PLAN BENEFITS

    Because employee awards under the Incentive Plans are discretionary, it is
not possible to determine the benefits or amounts that will be received in the
future by individual employees or groups of employees under the Incentive Plans.
Awards of Options to non-employee directors are also discretionary. However, it
is expected that each non-employee director will receive on the date of the
meeting as part of his or her regular compensation a grant of Options to
purchase shares of Applera-Applied Biosystems stock and shares of Applera-Celera
Genomics stock. During fiscal year 2002, each non-employee director received a
grant of Options to purchase 8,000 shares of Applera-Applied Biosystems stock
and 2,000 shares of Applera-Celera Genomics stock. In addition, each
non-employee director will receive on the date of the meeting Director Stock
Awards of up to 1,200 shares of Applera-Applied Biosystems stock and up to 300
shares of Applera-Celera Genomics stock. See "THE BOARD OF
DIRECTORS--COMPENSATION OF DIRECTORS," above.

                                       34

EQUITY COMPENSATION PLAN INFORMATION

    The following table sets forth information about shares of Applera common
stock that may be issued under the Company's equity compensation plans,
including compensation plans that were approved by the Company's stockholders as
well as compensation plans that were not approved by the Company's stockholders.
Information in the table is as of June 30, 2002.



                                                                                                 NUMBER OF SHARES REMAINING
                                    NUMBER OF SHARES TO BE       WEIGHTED-AVERAGE EXERCISE      AVAILABLE FOR FUTURE ISSUANCE
                                    ISSUED UPON EXERCISE OF        PRICE OF OUTSTANDING           UNDER EQUITY COMPENSATION
                                     OUTSTANDING OPTIONS,         OPTIONS, WARRANTS, AND         PLANS (EXCLUDING SECURITIES
                                     WARRANTS, AND RIGHTS                 RIGHTS                  REFLECTED IN COLUMN (A))
PLAN CATEGORY                                 (A)                           (B)                              (C)
-------------                       -----------------------      -------------------------      -----------------------------
                                                                                       
APPLERA-APPLIED BIOSYSTEMS STOCK

Equity compensation plans approved
by stockholders...................         33,741,457(1)                   $37.60                        10,357,019(2)

Equity compensation plans not
approved by stockholders..........                  0(3)

Total.............................         33,741,457                      $37.40                        10,357,019

APPLERA-CELERA GENOMICS STOCK

Equity compensation plans approved
by stockholders...................         10,433,176(4)                   $25.41                         4,432,626(5)

Equity compensation plans not
approved by stockholders..........            444,518(6,7)                 $30.15                           214,594(7)

Total.............................         10,877,694                      $25.61                         4,647,220


------------------------------

(1)   Represents shares of Applera-Applied Biosystems stock issuable upon the
      exercise of options outstanding under the following equity compensation
      plans: The Perkin-Elmer Corporation 1988 Stock Incentive Plan for Key
      Employees; The Perkin-Elmer Corporation 1993 Stock Incentive Plan for Key
      Employees; The Perkin-Elmer Corporation 1996 Stock Incentive Plan; The
      Perkin-Elmer Corporation 1997 Stock Incentive Plan; and The Perkin-Elmer
      Corporation 1998 Stock Incentive Plan (collectively, the "Frozen Applera
      Equity Plans"); and the Applera Corporation/Applied Biosystems Group 1999
      Stock Incentive Plan.

(2)   Represents shares of Applera-Applied Biosystems stock issuable pursuant to
      options and other rights authorized for future issuance under the Applera
      Corporation/Applied Biosystems Group 1999 Stock Incentive Plan and the
      Applera Corporation 1999 Employee Stock Purchase Plan.

(3)   As of June 30, 2002, options to purchase 299,007 shares of Applera-Applied
      Biosystems stock were outstanding under the following equity compensation
      plans, which options were assumed in connection with merger and
      acquisition transactions: the Applied Biosystems, Inc. 1992 Stock Option
      Plan; the Molecular Informatics, Inc. 1997 Equity Ownership Plan; the
      PerSeptive Biosystems 1992 Stock Plan; and the PerSeptive Biosystems 1997
      Non-Qualified Stock Option Plan. The weighted-average exercise price of
      these options as of June 30, 2002 was $13.98. No new options or other
      rights to equity compensation will be issued under these equity
      compensation plans, and the options outstanding under these equity
      compensation plans are not reflected in the table above.

(4)   Represents shares of Applera-Celera Genomics stock issuable upon the
      exercise of options outstanding under the Frozen Applera Equity Plans and
      the Applera Corporation/Celera Genomics Group 1999 Stock Incentive Plan.

(5)   Represents shares of Applera-Celera Genomics stock issuable pursuant to
      options and other rights authorized for future issuance under the Applera
      Corporation/Celera Genomics Group 1999 Stock Incentive Plan and the
      Applera Corporation 1999 Employee Stock Purchase Plan.

(6)   As of June 30, 2002, options to purchase 862,667 shares of Applera-Celera
      Genomics stock were outstanding under the following equity compensation
      plans, which options were assumed in connection with merger and
      acquisition transactions: the Applied Biosystems, Inc. 1992 Stock Option
      Plan; the Molecular Informatics, Inc. 1997 Equity Ownership Plan; the Axys
      Pharmaceuticals, Inc. 1989 Stock Option Plan; the Axys
      Pharmaceuticals, Inc. 1997 Equity Incentive Plan; the Axys
      Pharmaceuticals, Inc. 1997 Non-Officer Equity Incentive Plan; the
      PerSeptive Biosystems 1992 Stock Plan; the PerSeptive Biosystems 1997
      Non-Qualified Stock Option Plan; and the Paracel, Inc. Stock Option Plan.
      The weighted average exercise price of these options as of June 30, 2002
      was $25.22. No new options or other rights to equity compensation will be
      issued under these equity compensation plans, and the options outstanding
      under these equity compensation plans are not reflected in the table
      above, except for the Axys Pharmaceuticals, Inc. 1997 Equity Incentive
      Plan.

(7)   Represents shares of Applera-Celera Genomics stock issuable pursuant to
      options outstanding under the Axys Pharmaceuticals, Inc. 1997 Equity
      Incentive Plan, and shares of Applera-Celera Genomics stock issuable
      pursuant to options and other rights authorized for future issuance under
      that plan.

                                       35

    The following is a description of the material features of the Company's
equity compensation plans that were not approved by the Company's stockholders:

    APPLIED BIOSYSTEMS, INC. 1992 STOCK OPTION PLAN.  This plan was assumed by
the Company in connection with the acquisition of Applied Biosystems, Inc. No
new options or other rights to equity compensation will be issued under this
plan. As of June 30, 2002, there were options to purchase 5,052 shares of
Applera-Applied Biosystems stock and 1,302 shares of Applera-Celera Genomics
stock outstanding under this plan. The last of these options that were issued
are scheduled to terminate in October 2002.

    MOLECULAR INFORMATICS, INC. 1997 EQUITY OWNERSHIP PLAN.  This plan was
assumed by the Company in connection with the acquisition of Molecular
Informatics, Inc. No new options or other rights to equity compensation will be
issued under this plan. As of June 30, 2002, there were options to purchase
17,556 shares of Applera-Applied Biosystems stock and 3,342 shares of
Applera-Celera Genomics stock outstanding under this plan. The last of these
options that were issued are scheduled to terminate in July 2007.

    PERSEPTIVE BIOSYSTEMS 1992 STOCK PLAN.  This plan was assumed by the Company
in connection with the acquisition of PerSeptive Biosystems, Inc. No new options
or other rights to equity compensation will be issued under this plan. As of
June 30, 2002, there were options to purchase 260,483 shares of Applera-Applied
Biosystems stock and 67,880 shares of Applera-Celera Genomics stock outstanding
under this plan. The last of these options that were issued are scheduled to
terminate in July 2007.

    PERSEPTIVE BIOSYSTEMS 1997 NON-QUALIFIED STOCK OPTION PLAN.  This plan was
assumed by the Company in connection with the acquisition of PerSeptive
Biosystems, Inc. No new options or other rights to equity compensation will be
issued under this plan. As of June 30, 2002, there were options to purchase
15,916 shares of Applera-Applied Biosystems stock and 578 shares of
Applera-Celera Genomics stock outstanding under this plan. The last of these
options that were issued are scheduled to terminate in August 2007.

    PARACEL, INC. STOCK OPTION PLAN.  This plan was assumed by the Company in
connection with the acquisition of Paracel, Inc. No new options or other rights
to equity compensation will be issued under this plan. As of June 30, 2002,
there were options to purchase 178,992 shares of Applera-Celera Genomics stock
outstanding under this plan. The last of these options that were issued are
scheduled to terminate in February 2007.

    AXYS PHARMACEUTICALS, INC. 1989 STOCK OPTION PLAN.  This plan was assumed by
the Company in connection with the acquisition of Axys Pharmaceuticals, Inc. No
new options or other rights to equity compensation will be issued under this
plan. As of June 30, 2002, there were options to purchase 50,976 shares of
Applera-Celera Genomics stock outstanding under this plan. The last of these
options that were issued are scheduled to terminate in May 2009.

    AXYS PHARMACEUTICALS, INC. 1997 EQUITY INCENTIVE PLAN.  This plan was
assumed by the Company in connection with the acquisition of Axys
Pharmaceuticals, Inc. As of June 30, 2002, there were options to purchase
444,518 shares of Applera-Celera Genomics stock outstanding under this plan. The
last of these options that were issued are scheduled to terminate in
November 2011. 214,594 shares of Applera-Celera Genomics stock are authorized
for future issuance as equity compensation under this plan pursuant to stock
options, stock awards, and stock purchase awards. Employees and directors of and
consultants to Axys Pharmaceuticals, a wholly owned subsidiary of the Company,
and its affiliates are generally eligible for the grant of equity compensation
under this plan. The exercise price, vesting period, and all other terms and
conditions of each option granted under this plan will be determined by the
Company's Management Resources Committee, except that the exercise price may not
be less than the fair market value on the date of grant, and the term of each
option may not be more than 10 years. Stock awards and stock purchase awards
under this plan may be subject to such

                                       36

restrictions as may be determined by the Committee and may be subject to
repurchase rights in favor of the Company. Stock purchase awards under this plan
may not have a purchase price less than the fair market value on the date of the
award. This plan expires in November 2007, after which no equity compensation
may be issued under this plan.

    AXYS PHARMACEUTICALS, INC. 1997 NON-OFFICER EQUITY INCENTIVE PLAN.  This
plan was assumed by the company in connection with the acquisition of Axys
Pharmaceuticals, Inc. No new options or other rights to equity compensation will
be issued under this plan. As of June 30, 2002, there were options to purchase
115,079 shares of Applera-Celera Genomics stock outstanding under this plan. The
last of these options that were issued are scheduled to terminate in
October 2011.

VOTE REQUIRED FOR APPROVAL

    Approval of each proposal requires the favorable vote of a majority of votes
present in person or by proxy and entitled to vote at the meeting.

    THE BOARD RECOMMENDS THAT YOU VOTE "FOR" EACH OF THESE PROPOSALS.

                                 OTHER BUSINESS

    As of the date of this proxy statement, we do not know of any matter to be
brought before the meeting other than those described in this proxy statement.
If any other matters properly come before the meeting, the persons named as
proxies on the accompanying proxy card will vote on these matters in accordance
with their best judgment.

                             STOCKHOLDER PROPOSALS

    Any stockholder who wishes to submit a proposal to be included in the proxy
statement for the Company's 2003 annual meeting must deliver the proposal to the
Company no later than May 7, 2003. All proposals should be sent in writing to
the Secretary of the Company, 301 Merritt 7, P.O. Box 5435, Norwalk,
Connecticut, 06856-5435, and must include specified information about the
proposal and stockholder required by the SEC.

    The Company's By-laws also provide that any stockholder who intends to
present a nomination for a directorship or a proposal for action at any annual
meeting of stockholders must give advance notice of such proposal together with
certain specified information. These requirements are separate and apart from
and in addition to the SEC requirements noted above that a stockholder must meet
in order to have a proposal included in the Company's proxy materials. In
general, the advance notice must be given to the Secretary of the Company not
less than 45 days or more than 75 days prior to the first anniversary of the
date on which proxy materials for the preceding year's annual meeting were first
mailed to stockholders. In the case of the 2003 annual meeting, this advance
notice must be received no earlier than June 23, 2003 or later than July 23,
2003. The Company will have discretionary authority to vote on any stockholder
proposals presented at the 2003 annual meeting which do not comply with these
notice requirements. Additional information regarding the submission of
stockholder proposals may be obtained by writing to the Secretary of the Company
at the address provided above.

                                          By Order of the Board of Directors,
                                          Thomas P. Livingston
                                          SECRETARY

                                          Norwalk, Connecticut
                                          September 4, 2002

                                       37

                                   [GRAPHIC]



                                                    APPENDIX A TO SCHEDULE 14A
                                             FILED UNDER SCHEDULE 14A, ITEM 10


                               APPLERA CORPORATION

                        1999 EMPLOYEE STOCK PURCHASE PLAN

                           (AS PROPOSED TO BE AMENDED)


1.  PURPOSE OF THE PLAN.

         The purpose of the Applera Corporation 1999 Employee Stock Purchase
Plan (the "Plan") is to provide an incentive for Eligible Employees to continue
to devote their best efforts to the success of the Corporation, and to afford
such employees an opportunity to obtain a proprietary interest in the continued
growth and prosperity of the Corporation through ownership of its Common Stock
acquired in a convenient fashion.

2.  DEFINITIONS.

         As used herein, the following terms have the meanings hereinafter set
forth unless the context clearly indicates to the contrary:

         2.1 "ACT" means the Securities Exchange Act of 1934, as amended from
time to time.

         2.2 "APPLIED BIOSYSTEMS STOCK" means the Applera Corporation - Applied
Biosystems Group Common Stock, par value $.01 per share.

         2.3  "BOARD OF DIRECTORS" means the Board of Directors of the
Corporation.

         2.4 "CELERA STOCK" means the Applera Corporation - Celera Genomics
Group Common Stock, par value $.01 per share.

         2.5 "CODE" means the Internal Revenue Code of 1986, as amended from
time to time.

         2.6 "COMMITTEE" means the Management Resources Committee of the Board
of Directors, or any successor thereto or committee designated thereby.

         2.7 "COMMON STOCK" means Celera Stock and Applied Biosystems Stock and
either of them as the context requires.

         2.8 "COMPENSATION" means the regular basic wage or salary, including
commissions, paid to an Eligible Employee by the Corporation and any amount
which is contributed by the Corporation pursuant to a salary reduction agreement
and which is not includable in the gross income of the Eligible Employee under
Sections 125 and 402(g) of the Code or because it is





made to a deferred compensation plan sponsored by the Corporation. Bonus,
payment for overtime, or other special payments shall not be considered as part
of Compensation.

         2.9 "CORPORATION" means Applera Corporation and such of its
Subsidiaries existing as of the effective date of the Plan or thereafter
acquired as may be designated from time to time by the Committee.

         2.10 "DATE OF OFFERING" means the first date in the applicable Purchase
Period on which sales of Common Stock are made on a national securities exchange
unless another date is specified by the Committee.

         2.11 "ELIGIBLE EMPLOYEE" means any person who is an employee of the
Corporation on a Date of Offering during the term of the Plan. Directors of the
Corporation who are not employees and any employee who, immediately after the
grant of an option hereunder, would own (within the meaning of Section 424(d) of
the Code) Common Stock (including stock which such employee may purchase under
outstanding options) possessing 5% or more of the total combined voting power or
value of all classes of the capital stock of the Corporation or of a Subsidiary,
shall be ineligible to participate in the Plan.

         2.12 "FAIR MARKET VALUE" means the simple average of the high and low
sales prices of a share of Celera Stock or Applied Biosystems Stock, as the case
may be, as reported in the report of composite transactions (or other source
designated by the Committee) on the date on which fair market value is to be
determined (or if there shall be no trading on such date, then on the first
previous date on which sales were made on a national securities exchange).

         2.13 "OFFERING PRICE" means the lower of (a) 85% of the Fair Market
Value of a share of Celera Stock or Applied Biosystems Stock, as the case may
be, on the applicable Date of Offering, and (b) 85% of the Fair Market Value of
a share of such class of Common Stock on the last day of the applicable Purchase
Period on which sales of such class of Common Stock are made on a national
securities exchange unless another date is specified by the Committee.

         2.14 "PARTICIPATING EMPLOYEE" means an Eligible Employee who has
accepted all or any part of an option to purchase shares of Celera Stock,
Applied Biosystems Stock, or any combination thereof under an offering pursuant
to Section 7 hereof.

         2.15 "PURCHASE PERIOD" means each period of three calendar months
commencing on January 1, April 1, July 1, and October 1.

         2.16 "SUBSIDIARY" means any corporation in respect of which the
Corporation owns, directly or indirectly, more than 50% of the total combined
voting power of all classes of stock issued by such corporation.

3.  SHARES RESERVED FOR THE PLAN.

                                      -2-


         The aggregate number of shares of Applied Biosystems Stock available
for issuance under the Plan is Three Million (3,000,000), subject to adjustment
in accordance with Section 16 hereof. The aggregate number of shares of Celera
Stock available for issuance under the Plan is Two Million Seven Hundred
Thousand (2,700,000), subject to adjustment in accordance with Section 16
hereof. Shares of Common Stock issued under the Plan shall be authorized but
unissued shares. In lieu of such unissued shares, the Corporation may, in its
discretion, deliver treasury shares, reacquired shares, or shares acquired in
the market for purposes of the Plan.

         If any option granted under the Plan shall for any reason terminate, be
canceled, or expire without having been exercised, shares of Common Stock not
issued under such option shall be available again for issuance under the Plan.

4.  ADMINISTRATION OF THE PLAN.

         The Committee shall have plenary authority in its discretion, but
subject to the express provisions of the Plan, to administer the Plan. The
Committee shall also have plenary authority in its discretion to interpret the
Plan, to prescribe, amend, and rescind rules and regulations relating to it, and
to make any and all other determinations and take any and all actions deemed
necessary or advisable for the administration of the Plan. The Committee's
determination on the foregoing matters shall be conclusive and binding on all
persons having an interest in the Plan.

5.  OFFERINGS.

         Subject to the terms and conditions of the Plan, the Corporation may
make offerings to Eligible Employees to purchase shares of Common Stock under
the Plan during each of the five calendar years commencing January 1, 2000.

6.  AMOUNT OF COMMON STOCK EACH ELIGIBLE EMPLOYEE MAY PURCHASE.

         6.1 AMOUNT OF PURCHASE. Subject to the terms of the Plan, and as to
each offering made hereunder, each Eligible Employee shall be offered an option
to purchase that number of whole and fractional shares of Celera Stock and/or
Applied Biosystems Stock equal to (a) the total amount accumulated in such
Eligible Employee's account established pursuant to Section 8 hereof with
respect to such offering as of the last day of the applicable Purchase Period
divided by (b) the Offering Price of the Celera Stock or Applied Biosystems
Stock, as the case may be.

         6.2 LIMITATIONS ON PURCHASES. No Eligible Employee shall be granted an
option to purchase shares of Common Stock under all employee stock purchase
plans (to which Section 423 of the Code is applicable) of the Corporation and
its subsidiaries at a rate which exceeds $25,000 of the Fair Market Value of the
Common Stock (determined as of the date of grant of such option) for each
calendar year during which any option granted to such individual under any such
plan is outstanding at any time.

7.  METHOD OF PARTICIPATION.


                                      -3-


         7.1 NOTICE OF OFFERING. The Committee shall give notice to each
Eligible Employee of each offering under the Plan and the terms and conditions
of such offering.

         7.2 ELECTION BY ELIGIBLE EMPLOYEES. Each Eligible Employee who desires
to accept all or any part of the option to purchase shares of Common Stock under
an offering shall signify his or her election to do so in the form and manner
prescribed by the Committee. Each such Eligible Employee shall also authorize
the Corporation to make payroll deductions in accordance with Section 8 hereof
to cover the aggregate purchase price of those shares in respect of which he or
she has elected to accept an option. Such election and authorization shall
continue in effect for each subsequent offering unless at least ten (10) days
prior to the first day of the next succeeding Purchase Period the Eligible
Employee withdraws from the Plan or terminates employment with the Corporation,
as hereinafter provided, or elects a different rate of payroll deductions in the
form and manner prescribed by the Committee.

8.  PAYROLL DEDUCTIONS.

         8.1 PAYROLL DEDUCTIONS. Each Participating Employee shall authorize the
Corporation, in the form and manner prescribed by the Committee, to make payroll
deductions equal to any whole percentage of such Eligible Employee's
Compensation up to a maximum of 10% to cover the aggregate purchase price of
those shares in respect of which he or she has elected to accept an option.
Payroll deductions shall be deducted from such Participating Employee's
compensation through regular payroll deductions, and shall commence as soon as
practicable following the applicable Date of Offering and shall continue for the
duration of the Purchase Period. A separate bookkeeping account shall be
maintained by the Corporation for each Participating Employee, and the amount of
each Participating Employee's payroll deductions shall be credited to such
account.

         8.2 CONFLICTS WITH LAW. If any law, rule, or regulation applicable to
any Eligible Employee prohibits the use of payroll deductions for purposes of
the Plan, or if such deductions impair or hinder the operation of the Plan, an
alternative method of payment approved by the Committee may be substituted for
such Eligible Employee.

9.  EXERCISE OF OPTION AND PURCHASE OF SHARES.

         9.1 EXERCISE OF OPTION. Unless a Participating Employee has
subsequently withdrawn from the offering pursuant to Section 12 hereof, such
Participating Employee's option shall be deemed to have been automatically
exercised as of the last day of the applicable Purchase Period and become on
such date an irrevocable obligation to purchase shares of Common Stock in
accordance with the provisions of the Plan. The number of whole and fractional
shares of Celera Stock and/or Applied Biosystems Stock so purchased by each such
Participating Employee shall be determined by dividing (a) the amount
accumulated in such Participating Employee's account by payroll deductions with
respect to the offering of such class of Common Stock by (b) the Offering Price
of Celera Stock or Applied Biosystems Stock, as the case may be.


                                      -4-


         9.2 OVERSUBSCRIPTION. In the event that, with respect to any offering
hereunder, Participating Employees become entitled to purchase more shares of
Celera Stock or Applied Biosystems Stock than the number of shares of such class
of Common Stock then available for issuance under the Plan, the aggregate number
of shares of such class of Common Stock then available shall be apportioned
among Participating Employees on a pro rata basis in accordance with the number
of shares of such class of Common Stock actually subscribed for by each such
Participating Employee, except that subscriptions to purchase one share shall,
to the extent possible, be honored in full.

10.  ISSUANCE OF SHARES.

         All full and fractional shares of Common Stock purchased by a
Participating Employee under the Plan shall be issued in book entry form and
credited to an account established in such Participating Employee's name at a
stock brokerage or other financial services company designated by the Committee.
Alternatively, the Committee may, in its sole discretion, cause the Corporation
to issue a certificate to a Participating Employee for the number of whole
shares of Common Stock purchased by such Participating Employee. In such event,
the Corporation shall pay to such Participating Employee an amount in cash equal
to any fractional share multiplied by the Fair Market Value of a share of Common
Stock on the date as of which the payment is made.

11.  RIGHTS AS A STOCKHOLDER.

         No Participating Employee shall be entitled to any rights or privileges
of a stockholder of the Corporation, including the right to receive any
dividends which may be declared on shares of Common Stock, until such time as
the full purchase price of such Participating Employee's shares has been paid
and shares have been issued to or for the account of such Participating Employee
in accordance with Section 10 hereof.

12.  WITHDRAWALS.

         12.1 RIGHT TO WITHDRAWAL. No later than ten (10) days prior to the end
of the Purchase Period with respect to any offering, a Participating Employee
may, by filing an appropriate notice with the Committee, direct the Corporation
to (a) make no further deductions from his or her Compensation with respect to
such offering, or (b) cancel his or her entire option under such offering. Such
notice shall be irrevocable. As soon as practicable following receipt of such
notice, the Corporation shall cease all payroll deductions with respect to such
offering by such Participating Employee. If the employee has directed that
payroll deductions be discontinued, any sums theretofore deducted in respect of
the offering shall, subject to the provisions of Section 13 hereof, be retained
by the Corporation until the end of the applicable Purchase Period, at which
time there shall be issued to or for the account of the employee that number of
whole and fractional shares which can be purchased with the sum deducted. If the
employee has directed that his or her option be canceled, the Corporation shall,
as soon as practicable following receipt of such notice, refund in cash, without
interest, all amounts credited to the account of such employee with respect to
the applicable offering.


                                      -5-



         12.2 WAIVER OF WITHDRAWAL RIGHT. Notwithstanding the provisions of
Section 12.1 above, a Participating Employee may, at any time prior to the
expiration of any Purchase Period, irrevocably elect to waive both the right to
direct the Corporation to make no further deductions from such Participating
Employee's Compensation with respect to any option granted hereunder and the
right to cancel the entire option, which election shall be made by the filing of
an appropriate notice to such effect with the Committee. Upon the filing of such
a notice, such Participating Employee shall be irrevocably obligated to purchase
all of the shares of Common Stock covered by the option to which such notice
relates.

13.  TERMINATION OF EMPLOYMENT.

         13.1 DEATH, DISABILITY, OR RETIREMENT. In the event that the employment
of a Participating Employee is terminated prior to the end of a Purchase Period
because of total and permanent disability, retirement, or death, such
Participating Employee or his or her legal representative, as applicable, may
either:

                  (a) cancel his or her entire option with respect to such
         offering, in which event the Corporation shall, as soon as practicable
         thereafter, refund in cash, without interest, all amounts credited to
         such Participating Employee's account with respect to such offering; or

                  (b) elect to receive at the conclusion of the applicable
         Purchase Period that number of whole and fractional shares of Common
         Stock which such Participating Employee's payroll deductions actually
         made are sufficient to purchase.

         13.2 ELECTION. The election of a Participating Employee or his or her
legal representative, as applicable, pursuant to Section 13.1 above, shall be
made not later than ten (10) days prior to the end of the applicable Purchase
Period. Notification of the election shall be filed in the form and manner
prescribed by the Committee and, in the event that no notification has been
filed within the prescribed period, the Corporation shall act in accordance with
Section 13.1(a) above.

         13.3 OTHER TERMINATION OF EMPLOYMENT. In the event that the employment
of a Participating Employee is terminated for any reason other than those
specified in Section 13.1 above, the Corporation shall, as soon as practicable
thereafter, refund in cash, without interest, all amounts credited to such
Participating Employee's accounts under the Plan.

         13.4 TEMPORARY ABSENCE. In the event that the payroll deductions of a
Participating Employee are temporarily discontinued because of leave of absence,
temporary disability, or other similar reasons, then the number of shares of
Common Stock subject to purchase by such Participating Employee in any offering
shall be automatically reduced to that number of whole and fractional shares
which his or her aggregate payroll deductions actually made within the Purchase
Period are sufficient to purchase. Notwithstanding the foregoing, such
Participating Employee may make arrangements to pay to the Corporation an amount
equal to the amount which was not subject to payroll deductions by reason of the
temporary discontinuance thereof,

                                      -6-


and, in that event, such Participating Employee shall then be entitled to
purchase the total number of shares of Common Stock for which he or she has
accepted an option provided that full payment for all such shares is made not
later than the last day of the applicable Purchase Period.

14.  RIGHTS NOT TRANSFERABLE.

         A Participating Employee's rights under the Plan are exercisable,
during his or her lifetime, only by such Participating Employee and may not be
sold, pledged, assigned, or transferred in any manner. Any attempt to sell,
pledge, assign, or transfer such rights shall be void and unenforceable against
the Corporation or any affiliate. After the death of a Participating Employee,
such Participating Employee's rights may be transferred pursuant to the laws of
descent and distribution.

15.  NO RIGHT TO CONTINUED EMPLOYMENT.

         Nothing contained in the Plan shall confer upon any employee the right
to continue in the employ of the Corporation or any Subsidiary or interfere with
the right of the Corporation or such Subsidiary to terminate such employee's
employment at any time.

16.  ADJUSTMENT UPON CHANGES IN CAPITALIZATION.

         Notwithstanding any other provision of the Plan, in the event of
changes in the outstanding Celera Stock or Applied Biosystems Stock, as the case
may be, by reason of stock dividends, stock splits, recapitalizations,
combinations or exchanges of shares, corporate separations or divisions
(including, but not limited to, split-ups, split-offs, or spin-offs),
reorganizations (including, but not limited to, mergers or consolidations),
liquidations, or other similar events, the aggregate number and class of shares
available under the Plan and the number and class of shares under option but not
yet issued under the Plan shall be adjusted in such manner as the Committee in
its discretion deems appropriate.

17.  TERMINATION AND AMENDMENT OF THE PLAN.

         The Committee may terminate the Plan at any time or make such
modification or amendment to the Plan as it shall deem advisable. Upon
termination of the Plan, shares of Common Stock shall be issued to Participating
Employees as if the end of the applicable Purchase Period were the date of
termination of the Plan.

18.  GOVERNMENTAL REGULATIONS AND LISTING.

         All rights granted or to be granted to Eligible Employees under the
Plan are subject to all applicable laws and regulations and to the approval of
all governmental authorities required in connection with the authorization,
issuance, sale or transfer of the shares of Common Stock reserved for issuance
under the Plan, including, without limitation, there being a current
registration statement of the Corporation covering the offer of shares of Common
Stock purchasable under the options on the last day of the Purchase Period
applicable to such options,

                                      -7-


and if a registration statement shall not then be effective, the term of such
options and the Purchase Period shall be extended until the first business day
after the effective date of such registration statement, or post-effective
amendment thereto. In addition, all rights are subject to the due listing of
such shares of Common Stock on any securities exchange on which the Common Stock
is then listed.

         Notwithstanding any other provision of the Plan, the Plan is intended
to comply with all applicable provisions of Section 423 of the Code. To the
extent that any provision of the Plan or any action by the Committee under the
Plan fails to so comply, such provision or action shall, without further action
by any person, be deemed automatically amended to the extent necessary to effect
compliance with Section 423, provided that if such provision or action cannot be
amended to effect such compliance, such provision or action shall be deemed null
and void, to the extent permitted by law and deemed advisable by the Committee.
Each option granted to an Eligible Employee under the Plan shall be deemed
issued subject to the foregoing qualification.

19.  AWARDS IN FOREIGN COUNTRIES.

         The Committee shall have the authority and discretion to adopt such
modifications, procedures, and subplans as it shall deem necessary or desirable
to comply with the provisions of the laws of foreign countries in which the
Corporation may operate in order to assure the viability of the benefits of the
options made to individuals employed in such countries and to meet the
objectives of the Plan.

20.  GOVERNING LAW.

         Except where, and to the extent, offers of options under the Plan to
foreign employees are subject to foreign laws, the Plan shall be construed,
regulated, and administered under the internal laws of the State of Delaware.

21.  STOCKHOLDER APPROVAL.

         The Plan shall not become effective unless and until it has been
approved, in the manner prescribed by law, by the stockholders of the
Corporation.


                                      -8-




                                                     APPENDIX B TO SCHEDULE 14A
                                              FILED UNDER SCHEDULE 14A, ITEM 10


                  APPLERA CORPORATION/APPLIED BIOSYSTEMS GROUP

                            1999 STOCK INCENTIVE PLAN

                           (AS PROPOSED TO BE AMENDED)


1.  PURPOSE OF THE PLAN.

         The purpose of Applera Corporation/Applied Biosystems Group 1999 Stock
Incentive Plan (the "Plan") is to increase stockholder value and to advance the
interests of Applera Corporation and its subsidiaries (collectively, the
"Corporation") by providing financial incentives designed to attract, retain,
and motivate employees, officers, consultants, and directors of the Corporation.
The Plan continues the established policy of the Corporation of encouraging
ownership of its Stock by key personnel and of providing incentives for such
individuals to put forth maximum efforts for the success of the Corporation.

2.  DEFINITIONS.

         As used herein, the following terms have the meanings hereinafter set
forth unless the context clearly indicates to the contrary:

         2.1 "ACT" means the Securities Exchange Act of 1934, as amended from
time to time.

         2.2 "AGREEMENT" means the written agreement between the Corporation and
an Optionee or Award Recipient, as the case may be, evidencing the grant of an
Option or Award and setting forth the terms and conditions thereof.

         2.3  "AWARD" means a Stock Award or Performance Share Award.

         2.4 "AWARD RECIPIENT" means an individual to whom an Award has been
granted under the Plan.

         2.5  "BOARD OF DIRECTORS" means the Board of Directors of the
Corporation.

         2.6 "CODE" means the Internal Revenue Code of 1986, as amended from
time to time.

         2.7 "COMMITTEE" means the Management Resources Committee of the Board
of Directors, or any successor thereto or committee designated thereby whose
members




qualify as (a) outside directors as defined in Section 162(m) of the Code and
the Treasury Regulations issued pursuant thereto and (b) non-employee directors
within the meaning of Rule 16b-3 under the Act.

         2.8 "CONTINUOUS SERVICE" means an uninterrupted chain of continuous
regular employment by the Corporation or an uninterrupted chain of continuous
performance of significant services for the Corporation by a consultant. A leave
of absence granted in accordance with the Corporation's usual procedures which
does not operate to interrupt continuous employment or continuous performance of
significant services for other benefits granted by the Corporation shall not be
considered a termination of employment nor an interruption of Continuous Service
hereunder, and an employee or consultant who is granted such a leave of absence
shall be considered to be continuously employed or continuously performing
significant services during the period of such leave; PROVIDED, HOWEVER, that if
regulations under the Code or an amendment to the Code shall establish a more
restrictive definition of a leave of absence, such definition shall be
substituted herein.

         2.9 "DEFERRAL ACCOUNT" means the bookkeeping account established for
the deferral of a Director Stock Award by a Non-Employee Director pursuant to
Section 10.7 hereof.

         2.10 "DIRECTOR STOCK AWARD" means an award of shares of Stock granted
pursuant to Section 10 hereof.

         2.11  "EMPLOYEE AWARD" means an Employee Stock Award or Performance
Share Award.

         2.12 "EMPLOYEE STOCK AWARD" means an award of shares of Stock granted
pursuant to Section 8 hereof.

         2.13 "FAIR MARKET VALUE" means the simple average of the high and low
sales prices of a share of Stock as reported in the report of composite
transactions(or other source designated by the Committee) on the date on which
fair market value is to be determined (or if there shall be no trading on such
date, then on the first previous date on which sales were made on a national
securities exchange).

         2.14 "INCENTIVE STOCK OPTIONS" means those Options granted hereunder to
employees as incentive stock options as defined in, and which by their terms
comply with the requirements for such Options set out in, Section 422 of the
Code and the Treasury Regulations issued pursuant thereto.

         2.15 "NON-EMPLOYEE DIRECTOR" means a member of the Board of Directors
who is not an employee or officer of the Corporation.

         2.16 "NON-QUALIFIED STOCK OPTIONS" means those Options granted
hereunder which are not intended to qualify as Incentive Stock Options.

                                      -2-


         2.17 "NORMAL RETIREMENT AGE" means the normal retirement age of a
member of the Board as determined by the Board from time to time.

         2.18  "OPTION" means an option granted pursuant to Section 6 hereof.

         2.19 "OPTIONEE" means an individual to whom an Option has been granted
under the Plan.

         2.20 "PERFORMANCE SHARE AWARD" means an award of Performance Shares
granted pursuant to Section 9 hereof.

         2.21 "PERFORMANCE SHARES" means shares of Stock covered by a
Performance Share Award.

         2.22 "STOCK" means the Applera Corporation - Applied Biosystems Group
Common Stock, par value $.01 per share, of the Corporation.

         2.23  "STOCK AWARD" means an Employee Stock Award or Director Stock
Award.

         2.24 "STOCK UNIT" means the bookkeeping entry representing the
equivalent of one share of Stock.

         2.25 "STOCK RESTRICTIONS" mean the restrictions, including performance
goals, placed on a Stock Award or Performance Share Award under the Plan.

         2.26 "Ten Percent Stockholder" means an individual who owns, within the
meaning of Section 422(b)(6) of the Code and the Treasury Regulations issued
pursuant thereto, stock possessing more than ten (10%) percent of the total
combined voting power of all classes of stock of the Corporation.

3.  SHARES RESERVED FOR THE PLAN.

         The aggregate number of shares of Stock available for Options and
Awards under the Plan is 41,600,000, subject to adjustment in accordance with
Section 15. Shares of Stock issued under the Plan shall be authorized but
unissued shares. In lieu of such unissued shares, the Corporation may, in its
discretion, transfer on the exercise of Options or the delivery of shares of
Stock issued pursuant to Awards treasury shares, reacquired shares, or shares
acquired in the market for purposes of the Plan.

         If any Options or Awards granted under the Plan shall for any reason
terminate, be canceled, or expire without having been exercised or vested in
full, shares of Stock not issued or vested in full under such Options or Awards
shall be available again for issuance under the Plan.

4.  ADMINISTRATION OF THE PLAN.

                                      -3-


         The Committee shall have plenary authority in its discretion, but
subject to the express provisions of the Plan, to administer the Plan,
including, without limitation, the authority to determine the individuals to
whom, and the time or times at which, Options and Awards shall be granted, the
number of shares of Stock to be covered by each Option and Award, and the terms
and conditions of each Option and Award. The Committee shall also have plenary
authority in its discretion to interpret the Plan; to prescribe, amend, and
rescind rules and regulations relating to it; to determine the terms (which need
not be identical)of Agreements executed and delivered under the Plan, including,
without limitation, such terms and provisions as shall be requisite in the
judgment of the Committee to conform to any change in any law or regulation
applicable thereto; and to make any and all other determinations and take any
and all actions deemed necessary or advisable for the administration of the
Plan. The Committee's determination on the foregoing matters shall be conclusive
and binding on all persons having an interest in the Plan.

5.  ELIGIBILITY; FACTORS TO BE CONSIDERED IN GRANTING OPTIONS AND AWARDS.

         Subject to the terms of the Plan, an Option may be granted to any
person who, at the time the Option is granted, is a regular full-time employee
(which term shall include officers and directors) of the Corporation, a
Non-Employee Director, or a consultant performing significant services for the
Corporation. Employee Awards may be granted to any person who, at the time the
Employee Award is granted, is a regular full-time employee (which term shall
include officers and directors) of the Corporation. Non-Employee Directors shall
not be eligible to receive Employee Awards. In determining the employees,
Non-Employee Directors and consultants to whom Options or Awards shall be
granted, the number of shares of Stock to be covered by each Option or Award,
and the terms and conditions of each Option and Award, the Committee shall take
into account the duties and responsibilities of the respective employees,
Non-Employee Directors, and consultants, their present and potential
contributions to the success of the Corporation, and such other factors as they
shall deem relevant in connection with accomplishing the purposes of the Plan.
An employee, Non-Employee Director, or consultant who has been granted an Option
or Award may be granted and hold additional Options or Awards if the Committee
shall so determine.

6.  OPTIONS.

         6.1 GRANT OF OPTIONS. Subject to the terms of the Plan, the Committee
may grant Options to such employees, Non-Employee Directors, and consultants at
such time or times and in such amounts as it shall determine. Each Option
granted hereunder shall be designated as an Incentive Stock Option or
Non-Qualified Stock Option and shall be evidenced by an Agreement containing
such terms and conditions consistent with the Plan as the Committee shall
determine; PROVIDED, HOWEVER, that Incentive Stock Options shall be granted only
to employees of the Corporation.

                                      -4-


         6.2 PURCHASE PRICE. The purchase price of each share of Stock covered
by an Option shall be not less than 100% (or 110% in the case of an Incentive
Stock Option granted to a Ten Percent Stockholder) of the Fair Market Value of a
share of Stock on the date the Option is granted.

         6.3 TERM. The term of each Option shall be for such period as the
Committee shall determine, but not more than ten (10) years (or five (5) years
in the case of an Incentive Stock Option granted to a Ten Percent Stockholder)
from the date of grant thereof, and shall be subject to earlier termination as
hereinafter provided. If the original term of any Option is less than ten (10)
years (or five (5) years in the case of an Incentive Stock Option granted to a
Ten Percent Stockholder) from the date of grant, the Option prior to its
expiration may be amended, with the approval of the Committee and the employee,
Non-Employee Director, or consultant, as the case may be, to extend the term so
that the term as amended is not more than ten (10) years (or five (5) years in
the case of an Incentive Stock Option granted to a Ten Percent Stockholder) from
the original date of grant of such Option.

         6.4 VESTING. An Option shall be exercisable at such time or times and
in such manner and number of shares as the Committee shall determine. Except as
provided in the Plan, no Option may be exercised at any time unless the holder
thereof is then a regular employee of the Corporation, a member of the Board of
Directors, or a consultant performing significant services for the Corporation.
Options granted under the Plan shall not be affected by any change of duties or
position so long as the holder continues to be an employee of the Corporation,
continues to be a member of the Board of Directors, or a consultant performing
significant services for the Corporation.

         6.5 TERMINATION OF EMPLOYMENT OR SERVICES. Except as otherwise provided
in the Agreement, in the event that the employment of an employee to whom an
Option has been granted under the Plan shall be terminated or the services of a
Non-Employee Director or consultant to whom an Option has been granted under the
Plan shall be terminated (other than by reason of Cause, retirement, disability,
or death), such Option may, subject to the provisions of the Plan, be exercised,
to the extent that the employee, Non-Employee Director, or consultant was
entitled to do so at the date of termination of his or her employment or
services, at any time within thirty (30) days after such termination, but in no
event after the expiration of the term of the Option.

         6.6 TERMINATION OF EMPLOYMENT OR SERVICES FOR CAUSE. In the event that
the employment of an employee to whom an Option has been granted under the Plan
shall be terminated or the services of a Non-Employee Director or consultant to
whom an Option has been granted under the Plan shall be terminated for Cause (as
such term is defined below), such Option shall be immediately forfeited in full
upon such termination (regardless of the extent to which such Option may have
been exercisable as of such time). For purposes of this Section 6.6 only,
"Cause" shall be defined as (a) any act which is in bad faith and to the
detriment of the Corporation or (b) a material breach of any agreement with or
material obligation to the Corporation.

                                      -5-


         6.7 RETIREMENT OR DISABILITY. Except as otherwise provided in the
Agreement, if an employee to whom an Option has been granted under the Plan
shall retire from the Corporation pursuant to any qualified pension plan
provided by the Corporation, or if a Non-Employee Director (a) retires from the
Board of Directors upon reaching Normal Retirement Age or (b) resigns or
declines to stand for reelection with the approval of the Board of Directors, or
if an employee, Non-Employee Director, or consultant to whom an Option has been
granted becomes totally and permanently disabled, such Option may be exercised,
notwithstanding the provisions of Section 6.4, in full without regard to the
period of Continuous Service after the Option was granted at any time (a) in the
case of an employee holding an Incentive Stock Option, within three (3) months
after such retirement or disability, but in no event after the expiration of the
term of the Option or (b) in the case of a Non-Qualified Stock Option, within
one (1) year (three (3) years in the case of a Non-Employee Director) after such
retirement, disability, resignation, or declining, but in no event after the
expiration of the term of the Option.

         6.8 DEATH. If an employee, Non-Employee Director, or consultant to whom
an Option has been granted under the Plan shall die while employed by the
Corporation, serving as a member of the Board of Directors, or engaged to
perform services for the Corporation, such Option may be exercised to the extent
that the employee, Non-Employee Director, or consultant was entitled to do so at
the date of his or her death, by his or her executor or administrator or other
person at the time entitled by law to the employee's, Non-Employee Director's,
or consultant's rights under the Option, at any time within such period, not
exceeding one (1) year after his or her death, as shall be prescribed in the
Agreement, but in no event after the expiration of the term of the Option.

7.  TERMS AND CONDITIONS APPLICABLE TO OPTIONS.

         7.1 TRANSFERABILITY. During the lifetime of an Optionee, an Option
shall not be transferable, except pursuant to a domestic relations order;
PROVIDED, HOWEVER, that the Committee may, in its sole discretion, permit an
Optionee to transfer a Non-Qualified Stock Option to (a) a member of the
Optionee's immediate family, (b) a trust, the beneficiaries of which consist
exclusively of members of the Optionee's immediate family, or (c) a partnership,
the partners of which consist exclusively of members of the Optionee's immediate
family. After the death of an Optionee, an Option may be transferred pursuant to
the laws of descent and distribution.

         7.2 METHOD OF EXERCISE. An Option may be exercised by giving written
notice to the Corporation specifying the number of shares of Stock to be
purchased; PROVIDED that, except as otherwise provided by the Committee, an
Option may not be exercised as to fewer than 100 shares, or the remaining
exercisable shares covered by the Option if fewer than 100, at any one time. No
Option may be exercised with respect to a fractional share. The purchase price
of the shares as to which an Option shall be exercised shall be paid in full at
the time of exercise at the election of the holder of an Option (a) in cash or
currency of the United States of America, (b) by tendering to the Corporation
shares of Stock owned by such holder for at least six (6) months having a Fair
Market Value

                                      -6-



equal to the cash exercise price applicable to the purchase price of the shares
as to which the Option is being exercised, (c) a combination of cash and/or
previously owned shares of Stock valued at Fair Market Value, or (d) by payment
of such other consideration as the Committee shall from time to time determine.
For purposes of the immediately preceding sentence, Fair Market Value shall be
determined as of the business day immediately preceding the day on which the
Option is exercised. Notwithstanding the foregoing, the Committee shall have the
right to modify, amend, or cancel the provisions of clauses (b) and (c) above at
any time upon prior notice to the holders of Options.

         7.3 STOCKHOLDER RIGHTS. An Optionee shall have none of the rights of a
stockholder with respect to the shares subject to an Option until such shares
have been registered upon the exercise of the Option on the transfer books of
the Corporation in the name of such Optionee and then only to the extent that
any restrictions imposed thereon by the Committee shall have lapsed.

         7.4 NO LOANS. Neither the Corporation, any company with which it is
affiliated, nor any of their respective subsidiaries may directly or indirectly
lend money to any person for the purpose of assisting such person in acquiring
or carrying shares of Stock issued upon the exercise of an Option.

         7.5 CONDITIONS PRECEDENT TO EXERCISE. Notwithstanding any other
provision of the Plan, but subject to the provisions of Section 11, the exercise
of an Option following termination of employment or service shall be subject to
the satisfaction of the conditions precedent that the Optionee has not (a)
rendered services or engaged directly or indirectly in any business which in the
opinion of the Committee competes with or is in conflict with the interests of
the Corporation; PROVIDED, HOWEVER, that the ownership by an Optionee of 5% or
less of any class of securities of a publicly traded company shall not be deemed
to violate this clause or (b) violated any written agreement with the
Corporation, including, without limitation, any confidentiality agreement. An
Optionee's violation of clause (a) or (b) of the preceding sentence shall result
in the immediate forfeiture of any Options held by such Optionee.

         7.6 LIMITATIONS ON THE GRANT OF OPTIONS. No one individual may be
granted an Option or Options under the Plan during any fiscal year of the
Corporation for an aggregate number of shares of Stock which exceeds 10% of the
total number of shares reserved for issuance under the Plan. The aggregate Fair
Market Value of the Stock (determined as of the date the Option is granted) with
respect to which Incentive Stock Options granted under the Plan and all other
stock option plans of the Corporation (or any parent or subsidiary of the
Corporation) are exercisable for the first time by any specific individual
during any calendar year shall not exceed $100,000. No Incentive Stock Option
may be granted hereunder to an individual who immediately after such Option is
granted is a Ten Percent Stockholder unless (a) the Option price is at least
110% of the fair market value of such stock on the date of grant and (b) the
Option may not be exercised more than five (5) years after the date of grant.

8.  EMPLOYEE STOCK AWARDS.

                                      -7-


         8.1 GRANT OF EMPLOYEE STOCK AWARDS. Subject to the terms of the Plan,
the Committee may grant Employee Stock Awards to such employees at such time or
times and in such amounts as it shall determine. Shares of Stock issued pursuant
to Employee Stock Awards may, but need not, be subject to such restrictions as
may be established by the Committee at the time of the grant and reflected in an
Agreement.

         8.2 RESTRICTIONS ON EMPLOYEE STOCK AWARDS. Except as provided in the
Plan, any shares of Stock subject to an Employee Stock Award with respect to
which Stock Restrictions have not been satisfied shall be forfeited and all
rights of the employee to such Employee Stock Award shall terminate without any
payment of consideration by the Corporation. Except as set forth in Section 8.5,
a recipient of an Employee Stock Award subject to Stock Restrictions shall
forfeit such award in the event of the termination of his or her employment
during the period the shares are subject to Stock Restrictions.

         8.3 STOCKHOLDER RIGHTS. The recipient of an Employee Stock Award shall
be entitled to such rights of a stockholder with respect to the shares of Stock
issued pursuant to such Employee Stock Award as the Committee shall determine,
including the right to vote such shares of Stock, except that cash and stock
dividends with respect to such shares may, at the discretion of the Committee,
be either paid currently or withheld by the Corporation for the Award
Recipient's account, and interest may be accrued on the amount of cash dividends
withheld at a rate and subject to such terms as determined by the Committee.

         The Committee, in its discretion, may cause a legend or legends to be
placed on any certificate representing shares issued pursuant to Employee Stock
Awards, which legend or legends shall make appropriate reference to the Stock
Restrictions imposed thereon. The Committee may also in its discretion require
that certificates representing shares issued pursuant to Employee Stock Awards
remain in the physical custody of the Corporation or an escrow holder until any
or all of the Stock Restrictions imposed under the Plan have lapsed.

         8.4 NON-TRANSFERABILITY. Prior to the time Stock Restrictions lapse,
none of the shares of Stock issued pursuant to an Employee Stock Award may be
sold, assigned, bequeathed, transferred, pledged, hypothecated, or otherwise
disposed of in any way by the Award Recipient.

         8.5 LAPSE OF RESTRICTIONS. In the event of the termination of
employment of an Award Recipient, prior to the lapse of Stock Restrictions, by
reason of death, total and permanent disability, retirement, or resignation or
discharge from employment other than discharge for cause, the Committee may, in
its discretion, remove any Stock Restrictions on all or a portion of the Stock
subject to an Employee Stock Award.

         8.6 LIMITATIONS ON EMPLOYEE STOCK AWARDS. No employee may receive an
Employee Stock Award representing more than 160,000 shares of Stock during any
fiscal year of the Corporation, and the maximum number of shares of Stock which
may be

                                      -8-



issued to all employees pursuant to Employee Stock Awards under the Plan shall
be 320,000, subject in each case to adjustment in accordance with Section 15.


9.  PERFORMANCE SHARE AWARDS.

         9.1 GRANT OF PERFORMANCE SHARE AWARDS. Subject to the terms of the
Plan, the Committee may grant Performance Share Awards to such employees at such
time or times and in such amounts as it shall determine. Stock issued pursuant
to a Performance Share Award shall be subject to the attainment of performance
goals relating to one or more criteria within the meaning of Section 162(m) of
the Code and the Treasury Regulations issued pursuant thereto, including,
without limitation, stock price, market share, sales, earnings per share, return
on equity, costs, and cash flow, as determined by the Committee from time to
time. Any such objectives and the period in which such objectives are to be met
shall be determined by the Committee at the time of the grant and reflected in
an Agreement; PROVIDED, HOWEVER, that the period in which such objectives are to
be met shall be not less than one year. Each Performance Share Award shall also
be subject to such other restrictions as the Committee may determine.

         9.2 DELIVERY OF PERFORMANCE SHARES. Certificates representing
Performance Shares shall be registered in the Award Recipient's name but shall
remain in the physical custody of the Corporation until the Committee has
determined that the performance goals and other Stock Restrictions with respect
to such Performance Shares have been met.

         9.3 STOCKHOLDER RIGHTS. The recipient of a Performance Share Award
shall be entitled to such rights of a stockholder with respect to the
Performance Shares as the Committee shall determine, including the right to vote
such shares of Stock, except that cash and stock dividends with respect to the
Performance Shares may, at the discretion of the Committee, be either paid
currently or withheld by the Corporation for the Award Recipient's account, and
interest maybe accrued on the amount of cash dividends withheld at a rate and
subject to such terms as determined by the Committee.

         9.4 NON-TRANSFERABILITY. Prior to the time shares of Stock issued
pursuant to a Performance Share Award are delivered to an Award Recipient, none
of such shares may be sold, assigned, bequeathed, transferred, pledged,
hypothecated, or otherwise disposed of in any way by the Award Recipient.

         9.5 LAPSE OF RESTRICTIONS. In the event of the termination of
employment of an Award Recipient, prior to the lapse of Stock Restrictions, by
reason of death, total and permanent disability, retirement, or resignation or
discharge from employment other than discharge for cause, the Committee may, in
its discretion, remove any Stock Restrictions on all or a portion of a
Performance Share Award, or determine the performance objectives with respect to
all or a portion of a Performance Share Award to have been attained; PROVIDED,
HOWEVER, that the Committee shall not be entitled to exercise such discretion to
the extent that the ability to exercise such discretion would cause the


                                      -9-


Performance Share Award not to qualify as performance based compensation under
Section 162(m) of the Code.

         9.6 LIMITATIONS ON PERFORMANCE SHARE AWARDS. No employee may receive
Performance Share Awards representing more than 400,000 shares of Stock during
any fiscal year of the Corporation, and the maximum number of shares of Stock
which may be issued to all employees pursuant to Performance Share Awards under
the Plan shall be 1,600,000, subject in each case to adjustment in accordance
with Section 15.

10.  DIRECTOR STOCK AWARDS.

         10.1 GRANT OF DIRECTOR STOCK AWARDS. As of the date of each election or
reelection to the Board of Directors, each Non-Employee Director shall
automatically be granted a Director Stock Award with respect to 1,200 shares of
Stock, subject to adjustment in accordance with Section 15. Notwithstanding the
foregoing, each Non-Employee Director first elected to the Board of Directors on
a date other than the date of an annual meeting of stockholders shall be granted
that number of whole shares of Stock equal to the number of shares then subject
to a Director Stock Award (after giving effect to Section 10.8) multiplied by a
fraction, the numerator of which shall be the number of months remaining until
the anticipated date of the next annual meeting of stockholders, and the
denominator of which shall be 12. All Director Stock Awards shall be evidenced
by an agreement containing such terms and conditions consistent with the Plan as
the Committee shall determine.

         10.2 VESTING. Each Director Stock Award shall vest in full on the date
immediately preceding the first annual meeting of stockholders next following
the date of grant; PROVIDED, HOWEVER, that, except as provided in the Plan, the
recipient thereof continues to serve as a member of the Board of Directors as of
such date.

         10.3 FORFEITURE OF DIRECTOR STOCK AWARDS. Except as provided in the
Plan, a recipient of a Director Stock Award shall forfeit any unvested shares of
Stock subject to the Director Stock Award, and all rights of the Non-Employee
Director to such unvested shares shall terminate without payment of
consideration by the Corporation, upon the termination of his or her service as
a member of the Board of Directors.

         10.4 STOCKHOLDER RIGHTS. Except as provided in Sections 10.5 and 10.7,
a recipient of a Director Stock Award shall be entitled to all rights of a
stockholder with respect to the shares of Stock issued pursuant to the Director
Stock Award, including the right to receive dividends and to vote such shares of
Stock; PROVIDED, HOWEVER, that stock dividends paid with respect to such shares
shall be restricted to the same extent as the underlying shares of Stock issued
pursuant to the Director Stock Award.

         The Committee shall cause a legend or legends to be placed on any
certificate representing shares issued pursuant to a Director Stock Award, which
legend or legends shall make appropriate reference to the terms of the Director
Stock Award and the Plan. The Committee shall also require that certificates
representing shares issued pursuant to

                                      -10-



Director Stock Awards remain in the physical custody of the Corporation or an
escrow holder until such shares have vested in accordance with the terms of the
Plan.

         10.5 NON-TRANSFERABILITY. Prior to vesting, none of the shares of Stock
issued pursuant to a Director Stock Award may be sold, assigned, bequeathed,
transferred, pledged, hypothecated, or otherwise disposed of in any way by the
recipient thereof.

         10.6 TERMINATION OF SERVICE. If a Non-Employee Director to whom a
Director Stock Award has been granted shall cease to serve as a director as a
result of (a) his or her death, (b) retiring from the Board of Directors upon
reaching Normal Retirement Age, (c) becoming totally and permanently disabled,
or (d) resigning with the approval of the Board of Directors, all shares subject
to such Director Stock Award shall be vested in full, notwithstanding the
provisions of Section 10.2, as of the date of termination of service.

         10.7 DEFERRAL ELECTION. A Non-Employee Director may elect to defer
receipt of any Director Stock Award by filing the appropriate deferral form with
the Corporate Secretary on or before December 15th of the calendar year prior to
the calendar year in which such deferral is to be effective. Notwithstanding the
foregoing, any person elected as a Non-Employee Director for the first time
shall be permitted to make his or her first deferral election no later than
twenty (20) days after such election. In no event, however, shall any deferral
be permitted to the extent prohibited by applicable law. Deferrals shall be
subject to the following terms and conditions:

                  (a) A Non-Employee Director may elect to defer receipt of a
         Director Stock Award until (i) a specified date in the future, (ii)
         cessation of his or her service as a member of the Board of Directors,
         or (iii) the end of the calendar year in which cessation of his or her
         service as a member of the Board of Directors occurs.

                  (b) There shall be established a Deferral Account on the books
         of the Corporation for each Non-Employee Director electing to defer a
         Director Stock Award pursuant to this Section 10.7. Deferrals shall be
         credited to the Non-Employee Director's Deferral Account in Stock Units
         in the following manner: on the award date to which the deferral
         election applies, the amount deferred shall be converted into a number
         of Stock Units equal to the number of shares of Stock awarded that are
         subject to the deferral election. A Non-Employee Director shall not
         have any voting rights with respect to any Stock Units held in his or
         her Deferral Account.

                  (c) Whenever cash dividends are paid with respect to shares of
         Stock, each Non-Employee Director's Deferral Account shall be credited
         on the payment date of such dividend with additional Stock Units
         (including fractional units to the nearest one/one hundredth (1/100))
         equal in value to the amount of the cash dividend paid on a single
         share of Stock multiplied by the number of Stock Units (including
         fractional units) credited to his or her Deferral Account as of the
         date

                                      -11-


         of record for dividend purposes. For purposes of crediting dividends,
         the value of a Stock Unit shall be the Fair Market Value of a share
         of Stock as of the payment date of the dividend.

                  (d) The number of Stock Units credited to each Non-Employee
         Director's Deferral Account shall be appropriately adjusted in the same
         manner and to the same extent Director Stock Awards are adjusted and
         modified pursuant to Section 15. In the event of a transaction subject
         to Section 11, the Board of Directors shall have the authority to amend
         the Plan to provide for the conversion of Stock Units credited to
         Deferral Accounts into units equal to shares of stock of the resulting
         or acquiring company (or a related company), as appropriate, if such
         stock is publicly traded or, if not, into cash of equal value on the
         effective date of such transaction. If pursuant to the preceding
         sentence cash is credited to a Non-Employee Director's Deferral
         Account, interest shall be credited thereon from the date such cash is
         received to the date of distribution quarterly, at the end of each
         calendar quarter, at a rate per annum (computed on the basis of a
         360-day year and a 91-day quarter) equal to the prime rate announced
         publicly by Citibank, N.A. at the end of such calendar quarter. If
         units representing publicly traded stock of the resulting or acquired
         company (or a related company) are credited to a Non-Employee
         Director's Deferral Account, dividends shall be credited thereto in the
         same manner as dividends are credited on Stock Units credited to such
         Deferral Accounts.

                  (e) Subject to Section 10.7(g), distributions of a
         Non-Employee Director's Deferral Account under the Plan shall be made
         as follows:

                           (i) If a Non-Employee Director has elected to defer a
                  Director Stock Award to a specified date in the future,
                  payment shall be as of such date and shall be made or shall
                  commence, as the case may be, within thirty (30) days after
                  the date specified;

                           (ii) If a Non-Employee Director has elected to defer
                  a Director Stock Award until cessation of his or her service
                  as a member of the Board of Directors, payment shall be as of
                  the date of such cessation of service and shall be made or
                  shall commence, as the case may be, within thirty (30) days
                  after the cessation of the Non-Employee Director's service as
                  a director; and

                           (iii) If a Non-Employee Director has elected to defer
                  a Director Stock Award until the end of the calendar year in
                  which the cessation of his or her service as a member of the
                  Board of Directors occurs, payment shall be made as of
                  December 31st of such year and shall be made or commence, as
                  the case may be, on December 31st of such year.

                  (f) Notwithstanding any elections pursuant to Sections 10.7(a)
         and/or (g) hereof, in the event of the death of the Non-Employee
         Director prior to the

                                      -12-



         distribution of his or her Deferral Account, the balance credited to
         such Deferral Account as of the date of his or her death shall be paid,
         as soon as reasonably possible thereafter, in a single distribution to
         the Non-Employee Director's beneficiary or beneficiaries designated on
         such Non-Employee Director's deferral election form. If no such
         election or designation has been made, such amounts shall be payable to
         the Non-Employee Director's estate.

                  (g) A Non-Employee Director may elect to have his or her
         Deferral Account under the Plan paid in a single distribution or equal
         annual installments, not to exceed ten (10) annual installments. To the
         extent a Deferral Account is deemed invested in Stock Units, such Stock
         Units shall be converted to Stock on the distribution date as provided
         in Section 10.7(h). To the extent deemed invested in units of any other
         stock, such units shall similarly be converted and distributed in the
         form of stock. To the extent invested in a medium other than Stock
         Units or other units, each such distribution hereunder shall be in the
         medium credited to the Deferral Account.

                  (h) To the extent a Deferral Account is deemed invested in
         Stock Units, a single distribution shall consist of the number of whole
         shares of Stock equal to the number of Stock Units credited to the
         Non-Employee Director's Deferral Account on the date as of which the
         distribution occurs. Cash shall be paid to a Non-Employee Director in
         lieu of a fractional share, determined by reference to the Fair Market
         Value of a share of Stock on the date as of which the distribution
         occurs. In the event a Non-Employee Director has elected to receive
         annual installment payments, each such payment shall be determined as
         follows:

                           (i) To the extent his or her Deferral Account is
                  deemed to be invested in Stock Units, each such payment shall
                  consist of the number of whole shares of Stock equal to the
                  number of Stock Units (including fractional units) credited to
                  the Deferral Account on the date as of which the distribution
                  occurs, divided by the number of annual installments remaining
                  as of such distribution date. Cash shall be paid to
                  Non-Employee Directors in lieu of fractional shares,
                  determined by reference to the Fair Market Value of a share of
                  Stock on the date as of which the distribution occurs.

                           (ii) To the extent his or her Deferral Account has
                  been credited in cash, each such payment shall be calculated
                  by dividing the value on the date the distribution occurs of
                  that portion of the Non-Employee Director's Deferral Account
                  which is in cash by the number of annual installments
                  remaining as of such distribution date.

         10.8 LIMITATIONS ON DIRECTOR STOCK AWARDS. The Aggregate Value (as such
term is defined below) of all Director Stock Awards granted to a Non-Employee
Director under the Plan in any fiscal year, together with all Director Stock
Awards granted to such Non-Employee Director under the Applera
Corporation/Celera Genomics Group 1999

                                      -13-



Stock Incentive Plan during such year, may not exceed the Director Budgeted
Amount (as such term is defined below) for such year. In the event that the
Aggregate Value of such awards exceeds the Director Budgeted Amount, the
Committee shall allocate to each Non-Employee Director shares of Stock and
shares of Applera Corporation - Celera Genomics Group Common Stock ("Celera
Genomics Stock"), in the ratio, as near as may be practicable, of the number of
shares of Stock then outstanding to the number of shares of Celera Genomics
Stock then outstanding, as determined by the Committee. For purposes of the
Plan, "Aggregate Value" shall be calculated as the Fair Market Value of a share
of Stock or Celera Genomics Stock, as the case may be, on the date of grant
multiplied by the number of shares of Stock or Celera Genomics Stock, as the
case may be, subject to the Director Stock Award, and the "Director Budgeted
Amount" shall be $135,000 for the fiscal year ending June 30, 2001 and shall
increase by 10% for each fiscal year thereafter.

11.  ACCELERATION UPON A CHANGE OF CONTROL.

         Notwithstanding any other provision of the Plan or any Option or Award
granted hereunder, (a) any Option granted hereunder and then outstanding shall
become immediately exercisable in full, (b) all Stock Restrictions shall
immediately terminate, and (c) all performance objectives applicable to any
Performance Share Award shall be deemed attained (i) in the event that a tender
offer or exchange offer (other than an offer by the Corporation) for common
stock of the Company representing more than 25% of the combined voting power of
the then outstanding voting securities of the Company entitled to vote generally
in the election of directors ("Voting Securities") is made by any "person"
within the meaning of Section 14(d) of the Act and not withdrawn within ten (10)
days after the commencement thereof; PROVIDED, HOWEVER, that the Committee may
by action taken prior to the end of such ten (10) day period extend such ten
(10) day period; and, PROVIDED FURTHER, that the Committee may by further action
taken prior to the end of such extended period declare (a) all Options granted
hereunder and then outstanding to be immediately exercisable in full, (b) all
Stock Restrictions to be immediately terminated, and (c) all performance
objectives applicable to any Performance Share Award to be deemed attained; or
(ii) in the event of a Change in Control (as hereinafter defined).

         For purposes of this Section 11, a "Change in Control" means an event
that would be required to be reported (assuming such event has not been
"previously reported") in response to Item 1(a) of the Current Report on Form
8-K, as in effect on the effective date of the Plan, pursuant to Section 13 or
15(d) of the Act; PROVIDED, HOWEVER, that, without limitation, such a Change in
Control shall be deemed to have occurred at such time as (a) any "person" within
the meaning of Section 14(d) of the Act becomes the "beneficial owner" as
defined in Rule 13d-3 thereunder, directly or indirectly, of more than 25% of
the combined voting power of the then outstanding Voting Securities, (b) during
any two-year period, individuals who constitute the Board of Directors (the
"Incumbent Board") as of the beginning of the period cease for any reason to
constitute at least a majority thereof, provided that any person becoming a
director during such period whose election or nomination for election by the
Corporation's stockholders was

                                      -14-



approved by a vote of at least three-quarters of the Incumbent Board (either by
a specific vote or by approval of the proxy statement of the Corporation in
which such person is named as a nominee for director without objection to such
nomination, other than in response to an actual or threatened Change in Control
or proxy contest) shall be, for purposes of this clause (b),considered as though
such person were a member of the Incumbent Board, or (c) the approval by the
Corporation's stockholders of the sale of all or substantially all of the stock
or assets of the Corporation. The Committee may adopt such procedures as to
notice and exercise as may be necessary to effectuate the acceleration of the
exercisability of Options, termination of Stock Restrictions, and attainment of
performance objectives as described above.

12.  SHARE WITHHOLDING.

         With respect to any Option or Award, the Committee may, in its
discretion and subject to such rules as the Committee may adopt, permit or
require any Optionee or Award Recipient to satisfy, in whole or in part, any
withholding tax obligation which may arise in connection with an Option or Award
by electing to have the Corporation withhold Stock having a Fair Market Value
(as of the date the amount of withholding tax is determined) equal to the amount
of withholding tax.

13.  NO RIGHT TO CONTINUED EMPLOYMENT OR SERVICE.

         Nothing contained in the Plan or in any Option or Award granted or
Agreement entered into pursuant to the Plan shall confer upon any employee the
right to continue in the employ of the Corporation, any consultant the right to
continue to perform services for the Corporation, or any Non-Employee Director
the right to continue as a member of the Board of Directors or interfere with
the right of the Corporation to terminate such employee's employment, such
consultant's service, or Non-Employee Director's service at any time.

14.  TIME OF GRANTING OPTIONS AND EMPLOYEE AWARDS.

         Nothing contained in the Plan or in any resolution adopted by the Board
of Directors or the holders of Stock shall constitute the grant of any Option or
Award hereunder. An Option or Award under the Plan shall be deemed to have been
granted on the date on which the name of the recipient and the terms of the
Option or Award are set forth in an Agreement and delivered to the recipient,
unless otherwise provided in the Agreement.

15.  ADJUSTMENTS UPON CHANGES IN CAPITALIZATION.

         Notwithstanding any other provision of the Plan, in the event of
changes in the outstanding Stock by reason of stock dividends, stock splits,
recapitalizations, combinations or exchanges of shares, corporate separations
or divisions (including, but not limited to, split-ups, split-offs, or
spin-offs), reorganizations (including, but not limited to, mergers or
consolidations),liquidations, or other similar events, the aggregate

                                      -15-


number and class of shares available under the Plan, the number of shares
subject to Director Stock Awards, the maximum number of shares that may be
subject to Options and Awards, and the terms of any outstanding Options or
Awards (including, without limitation, the number of shares subject to an
outstanding Option or Award and the price at which shares of Stock may be issued
pursuant to an outstanding Option) and of any Stock Units shall be adjusted in
such manner as the Committee in its discretion deems appropriate.

16.  TERMINATION AND AMENDMENT OF THE PLAN.

         Unless the Plan shall have been terminated as hereinafter provided, no
Option or Award shall be granted hereunder after March 31, 2004. The Board of
Directors may at any time prior to that date terminate the Plan or make such
modification or amendment to the Plan as it shall deem advisable; PROVIDED,
HOWEVER, that, except as provided in Section 15, no amendment may be made
without the approval by the holders of Stock (to the extent such approval would
be required for an exemption under Section 16(b) of the Act which the Company
wishes to have) if such amendment would (a) increase the aggregate number of
shares of Stock which may be issued under the Plan, (b) materially modify the
requirements as to eligibility for participation in the Plan, or (c) materially
increase the benefits accruing to participants under the Plan. No termination,
modification, or amendment of the Plan may, without the consent of an Optionee
or Award Recipient, adversely affect in any material manner the rights of such
Optionee or Award Recipient under any Option or Award.

17.  AMENDMENT OF OPTIONS AND AWARDS AT THE DISCRETION OF THE COMMITTEE.

         The terms of any outstanding Option or Award may be amended from time
to time by the Committee in its discretion in any manner that it deems
appropriate, including, without limitation, acceleration of the date of exercise
of any Option or Award, termination of Stock Restrictions as to any Award, or
the conversion of an Incentive Stock Option into a Non-Qualified Stock Option;
PROVIDED, HOWEVER, that no such amendment shall adversely affect in any material
manner any right of any Optionee or Award Recipient under the Plan without his
or her consent; and, PROVIDED FURTHER, that the Committee shall not (a) amend
any previously-issued Performance Share Award to the extent that such amendment
would cause such Performance Share Award not to qualify as performance based
compensation under Section 162(m) of the Code or (b) amend any previously-issued
Option to reduce the purchase price thereof whether by modification of the
Option or by cancellation of the Option in consideration of the immediate
issuance of a replacement Option bearing a reduced purchase price.

18.  GOVERNMENT REGULATIONS.

         The Plan and the grant and exercise of Options and Awards hereunder,
and the obligation of the Corporation to issue, sell, and deliver shares, as
applicable, under such Options and Awards, shall be subject to all applicable
laws, rules, and regulations. Notwithstanding any other provision of the Plan,
transactions under the Plan are intended

                                      -16-



to comply with the applicable exemptions under Rule 16b-3 under the Act as to
persons subject to the reporting requirements of Section 16(a) of the Act with
respect to shares of Stock, and Options and Awards under the Plan shall be
fashioned and administered in a manner consistent with the conditions applicable
under Rule 16b-3.

19.  OPTIONS AND AWARDS IN FOREIGN COUNTRIES.

         The Committee shall have the authority and discretion to adopt such
modifications, procedures, and subplans as it shall deem necessary or desirable
to comply with the provisions of the laws of foreign countries in which the
Corporation may operate in order to assure the viability of the benefits of the
Options and Awards made to individuals employed in such countries and to meet
the objectives of the Plan.

20.  GOVERNING LAW.

         The Plan shall be construed, regulated, and administered under the
internal laws of the State of Delaware.

21.  STOCKHOLDER APPROVAL.

         The Plan shall become effective upon the date of adoption by the Board
of Directors, subject to approval by the stockholders of the Corporation in
accordance with applicable law. Unless so approved within one (1) year after the
date of the adoption of the Plan by the Board of Directors, the Plan shall not
be effective for any purpose. Prior to approval by the Corporation's
stockholders, the Committee may grant Options and Awards under the terms of the
Plan, but if stockholder approval is not obtained in the specified period, such
Options and Awards shall be of no effect.


                                      -17-




                                                     APPENDIX C TO SCHEDULE 14A
                                              FILED UNDER SCHEDULE 14A, ITEM 10


                    APPLERA CORPORATION/CELERA GENOMICS GROUP

                            1999 STOCK INCENTIVE PLAN

                           (AS PROPOSED TO BE AMENDED)


1.  PURPOSE OF THE PLAN.

         The purpose of Applera Corporation/Celera Genomics Group 1999 Stock
Incentive Plan (the "Plan") is to increase stockholder value and to advance the
interests of Applera Corporation and its subsidiaries (collectively, the
"Corporation") by providing financial incentives designed to attract, retain,
and motivate employees, officers, consultants, and directors of the Corporation.
The Plan continues the established policy of the Corporation of encouraging
ownership of its Stock by key personnel and of providing incentives for such
individuals to put forth maximum efforts for the success of the Corporation.

2.  DEFINITIONS.

         As used herein, the following terms have the meanings hereinafter set
forth unless the context clearly indicates to the contrary:

         2.1 "ACT" means the Securities Exchange Act of 1934, as amended from
time to time.

         2.2 "AGREEMENT" means the written agreement between the Corporation and
an Optionee or Award Recipient, as the case may be, evidencing the grant of an
Option or Award and setting forth the terms and conditions thereof.

         2.3  "AWARD" means a Stock Award or Performance Share Award.

         2.4 "AWARD RECIPIENT" means an individual to whom an Award has been
granted under the Plan.

         2.5  "BOARD OF DIRECTORS" means the Board of Directors of the
Corporation.

         2.6 "CODE" means the Internal Revenue Code of 1986, as amended from
time to time.

         2.7 "COMMITTEE" means the Management Resources Committee of the Board
of Directors, or any successor thereto or committee designated thereby whose
members




qualify as (a) outside directors as defined in Section 162(m) of the Code and
the Treasury Regulations issued pursuant thereto and (b) non-employee directors
within the meaning of Rule 16b-3 under the Act.

         2.8 "CONTINUOUS SERVICE" means an uninterrupted chain of continuous
regular employment by the Corporation or an uninterrupted chain of continuous
performance of significant services for the Corporation by a consultant. A leave
of absence granted in accordance with the Corporation's usual procedures which
does not operate to interrupt continuous employment or continuous performance of
significant services for other benefits granted by the Corporation shall not be
considered a termination of employment nor an interruption of Continuous Service
hereunder, and an employee or consultant who is granted such a leave of absence
shall be considered to be continuously employed or continuously performing
significant services during the period of such leave; PROVIDED, HOWEVER, that if
regulations under the Code or an amendment to the Code shall establish a more
restrictive definition of a leave of absence, such definition shall be
substituted herein.

         2.9 "DEFERRAL ACCOUNT" means the bookkeeping account established for
the deferral of a Director Stock Award by a Non-Employee Director pursuant to
Section 10.7 hereof.

         2.10 "DIRECTOR STOCK AWARD" means an award of shares of Stock granted
pursuant to Section 10 hereof.

         2.11 "EMPLOYEE AWARD" means an Employee Stock Award or Performance
Share Award.

         2.12 "EMPLOYEE STOCK AWARD" means an award of shares of Stock granted
pursuant to Section 8 hereof.

         2.13 "FAIR MARKET VALUE" means the simple average of the high and low
sales prices of a share of Stock as reported in the report of composite
transactions(or other source designated by the Committee) on the date on which
fair market value is to be determined (or if there shall be no trading on such
date, then on the first previous date on which sales were made on a national
securities exchange).

         2.14 "INCENTIVE STOCK OPTIONS" means those Options granted hereunder to
employees as incentive stock options as defined in, and which by their terms
comply with the requirements for such Options set out in, Section 422 of the
Code and the Treasury Regulations issued pursuant thereto.

         2.15 "NON-EMPLOYEE DIRECTOR" means a member of the Board of Directors
who is not an employee or officer of the Corporation.

         2.16 "NON-QUALIFIED STOCK OPTIONS" means those Options granted
hereunder which are not intended to qualify as Incentive Stock Options.


                                      -2-



         2.17 "NORMAL RETIREMENT AGE" means the normal retirement age of a
member of the Board as determined by the Board from time to time.

         2.18  "OPTION" means an option granted pursuant to Section 6 hereof.

         2.19 "OPTIONEE" means an individual to whom an Option has been granted
under the Plan.

         2.20 "PERFORMANCE SHARE AWARD" means an award of Performance Shares
granted pursuant to Section 9 hereof.

         2.21 "PERFORMANCE SHARES" means shares of Stock covered by a
Performance Share Award.

         2.22 "STOCK" means the Applera Corporation - Celera Genomics Group
Common Stock, par value $.01 per share, of the Corporation.

         2.23  "STOCK AWARD" means an Employee Stock Award or Director
Stock Award.

         2.24 "STOCK UNIT" means the bookkeeping entry representing the
equivalent of one share of Stock.

         2.25 "STOCK RESTRICTIONS" mean the restrictions, including performance
goals, placed on a Stock Award or Performance Share Award under the Plan.

         2.26 "TEN PERCENT STOCKHOLDER" means an individual who owns, within the
meaning of Section 422(b)(6) of the Code and the Treasury Regulations issued
pursuant thereto, stock possessing more than ten (10%) percent of the total
combined voting power of all classes of stock of the Corporation.

3.  SHARES RESERVED FOR THE PLAN.

         The aggregate number of shares of Stock available for Options and
Awards under the Plan is 18,700,000, subject to adjustment in accordance with
Section 15. Shares of Stock issued under the Plan shall be authorized but
unissued shares. In lieu of such unissued shares, the Corporation may, in its
discretion, transfer on the exercise of Options or the delivery of shares of
Stock issued pursuant to Awards treasury shares, reacquired shares, or shares
acquired in the market for purposes of the Plan.

         If any Options or Awards granted under the Plan shall for any reason
terminate, be canceled, or expire without having been exercised or vested in
full, shares of Stock not issued or vested in full under such Options or Awards
shall be available again for issuance under the Plan.

4.  ADMINISTRATION OF THE PLAN.


                                      -3-



         The Committee shall have plenary authority in its discretion, but
subject to the express provisions of the Plan, to administer the Plan,
including, without limitation, the authority to determine the individuals to
whom, and the time or times at which, Options and Awards shall be granted, the
number of shares of Stock to be covered by each Option and Award, and the terms
and conditions of each Option and Award. The Committee shall also have plenary
authority in its discretion to interpret the Plan; to prescribe, amend, and
rescind rules and regulations relating to it; to determine the terms (which need
not be identical)of Agreements executed and delivered under the Plan, including,
without limitation, such terms and provisions as shall be requisite in the
judgment of the Committee to conform to any change in any law or regulation
applicable thereto; and to make any and all other determinations and take any
and all actions deemed necessary or advisable for the administration of the
Plan. The Committee's determination on the foregoing matters shall be conclusive
and binding on all persons having an interest in the Plan.

5.  ELIGIBILITY; FACTORS TO BE CONSIDERED IN GRANTING OPTIONS AND AWARDS.

         Subject to the terms of the Plan, an Option may be granted to any
person who, at the time the Option is granted, is a regular full-time employee
(which term shall include officers and directors) of the Corporation, a
Non-Employee Director, or a consultant performing significant services for the
Corporation. Employee Awards may be granted to any person who, at the time the
Employee Award is granted, is a regular full-time employee (which term shall
include officers and directors) of the Corporation. Non-Employee Directors shall
not be eligible to receive Employee Awards. In determining the employees,
Non-Employee Directors and consultants to whom Options or Awards shall be
granted, the number of shares of Stock to be covered by each Option or Award,
and the terms and conditions of each Option and Award, the Committee shall take
into account the duties and responsibilities of the respective employees,
Non-Employee Directors, and consultants, their present and potential
contributions to the success of the Corporation, and such other factors as they
shall deem relevant in connection with accomplishing the purposes of the Plan.
An employee, Non-Employee Director, or consultant who has been granted an Option
or Award may be granted and hold additional Options or Awards if the Committee
shall so determine.

6.  OPTIONS.

         6.1 GRANT OF OPTIONS. Subject to the terms of the Plan, the Committee
may grant Options to such employees, Non-Employee Directors, and consultants at
such time or times and in such amounts as it shall determine. Each Option
granted hereunder shall be designated as an Incentive Stock Option or
Non-Qualified Stock Option and shall be evidenced by an Agreement containing
such terms and conditions consistent with the Plan as the Committee shall
determine; PROVIDED, HOWEVER, that Incentive Stock Options shall be granted only
to employees of the Corporation.


                                      -4-


         6.2 PURCHASE PRICE. The purchase price of each share of Stock covered
by an Option shall be not less than 100% (or 110% in the case of an Incentive
Stock Option granted to a Ten Percent Stockholder) of the Fair Market Value of a
share of Stock on the date the Option is granted.

         6.3 TERM. The term of each Option shall be for such period as the
Committee shall determine, but not more than ten (10) years (or five (5) years
in the case of an Incentive Stock Option granted to a Ten Percent Stockholder)
from the date of grant thereof, and shall be subject to earlier termination as
hereinafter provided. If the original term of any Option is less than ten (10)
years (or five (5) years in the case of an Incentive Stock Option granted to a
Ten Percent Stockholder) from the date of grant, the Option prior to its
expiration may be amended, with the approval of the Committee and the employee,
Non-Employee Director, or consultant, as the case may be, to extend the term so
that the term as amended is not more than ten (10) years (or five (5) years in
the case of an Incentive Stock Option granted to a Ten Percent Stockholder) from
the original date of grant of such Option.

         6.4 VESTING. An Option shall be exercisable at such time or times and
in such manner and number of shares as the Committee shall determine. Except as
provided in the Plan, no Option may be exercised at any time unless the holder
thereof is then a regular employee of the Corporation, a member of the Board of
Directors, or a consultant performing significant services for the Corporation.
Options granted under the Plan shall not be affected by any change of duties or
position so long as the holder continues to be an employee of the Corporation,
continues to be a member of the Board of Directors, or a consultant performing
significant services for the Corporation.

         6.5 TERMINATION OF EMPLOYMENT OR SERVICES. Except as otherwise provided
in the Agreement, in the event that the employment of an employee to whom an
Option has been granted under the Plan shall be terminated or the services of a
Non-Employee Director or consultant to whom an Option has been granted under the
Plan shall be terminated (other than by reason of Cause, retirement, disability,
or death) such Option may, subject to the provisions of the Plan, be exercised,
to the extent that the employee, Non-Employee Director, or consultant was
entitled to do so at the date of termination of his or her employment or
services, at any time within thirty (30) days after such termination, but in no
event after the expiration of the term of the Option.

         6.6 TERMINATION OF EMPLOYMENT OR SERVICES FOR CAUSE. In the event that
the employment of an employee to whom an Option has been granted under the Plan
shall be terminated or the service of a Non-Employee Director or consultant to
whom an Option has been granted under the Plan shall be terminated for Cause (as
such term is defined below), such Option shall be immediately forfeited in full
upon such termination (regardless of the extent to which such Option may have
been exercisable as of such time). For purposes of this Section 6.6 only,
"Cause" shall be defined as (a) any act which is in bad faith and to the
detriment of the Corporation or (b) a material breach of any agreement with or
material obligation to the Corporation.


                                      -5-


         6.7 RETIREMENT OR DISABILITY. Except as otherwise provided in the
Agreement, if an employee to whom an Option has been granted under the Plan
shall retire from the Corporation pursuant to any qualified pension plan
provided by the Corporation, or if a Non-Employee Director (a) retires from the
Board of Directors upon reaching Normal Retirement Age or (b) resigns or
declines to stand for reelection with the approval of the Board of Directors, or
if an employee, Non-Employee Director, or consultant to whom an Option has been
granted becomes totally and permanently disabled, such Option may be exercised,
notwithstanding the provisions of Section 6.4, in full without regard to the
period of Continuous Service after the Option was granted at any time (a) in the
case of an employee holding an Incentive Stock Option, within three (3) months
after such retirement or disability, but in no event after the expiration of the
term of the Option or (b) in the case of a Non-Qualified Stock Option, within
one (1) year (three (3) years in the case of a Non-Employee Director) after such
retirement, disability, resignation, or declining, but in no event after the
expiration of the term of the Option.

         6.8 DEATH. If an employee, Non-Employee Director, or consultant to whom
an Option has been granted under the Plan shall die while employed by the
Corporation, serving as a member of the Board of Directors, or engaged to
perform services for the Corporation, such Option may be exercised to the extent
that the employee, Non-Employee Director, or consultant was entitled to do so at
the date of his or her death, by his or her executor or administrator or other
person at the time entitled by law to the employee's, Non-Employee Director's,
or consultant's rights under the Option, at any time within such period, not
exceeding one (1) year after his or her death, as shall be prescribed in the
Agreement, but in no event after the expiration of the term of the Option.

7.  TERMS AND CONDITIONS APPLICABLE TO OPTIONS.

         7.1 TRANSFERABILITY. During the lifetime of an Optionee, an Option
shall not be transferable, except pursuant to a domestic relations order;
PROVIDED, HOWEVER, that the Committee may, in its sole discretion, permit an
Optionee to transfer a Non-Qualified Stock Option to (a) a member of the
Optionee's immediate family, (b) a trust, the beneficiaries of which consist
exclusively of members of the Optionee's immediate family, or (c) a partnership,
the partners of which consist exclusively of members of the Optionee's immediate
family. After the death of an Optionee, an Option may be transferred pursuant to
the laws of descent and distribution.

         7.2 METHOD OF EXERCISE. An Option may be exercised by giving written
notice to the Corporation specifying the number of shares of Stock to be
purchased; PROVIDED that, except as otherwise provided by the Committee, an
Option may not be exercised as to fewer than 100 shares, or the remaining
exercisable shares covered by the Option if fewer than 100, at any one time. No
Option may be exercised with respect to a fractional share. The purchase price
of the shares as to which an Option shall be exercised shall be paid in full at
the time of exercise at the election of the holder of an Option (a) in cash or
currency of the United States of America, (b) by tendering to the Corporation
shares of Stock owned by such holder for at least six (6) months having a Fair
Market Value equal


                                      -6-



to the cash exercise price applicable to the purchase price of the shares as to
which the Option is being exercised, (c) a combination of cash and/or previously
owned shares of Stock valued at Fair Market Value, or (d) by payment of such
other consideration as the Committee shall from time to time determine. For
purposes of the immediately preceding sentence, Fair Market Value shall be
determined as of the business day immediately preceding the day on which the
Option is exercised. Notwithstanding the foregoing, the Committee shall have the
right to modify, amend, or cancel the provisions of clauses (b) and (c) above at
any time upon prior notice to the holders of Options.

         7.3 STOCKHOLDER RIGHTS. An Optionee shall have none of the rights of a
stockholder with respect to the shares subject to an Option until such shares
have been registered upon the exercise of the Option on the transfer books of
the Corporation in the name of such Optionee and then only to the extent that
any restrictions imposed thereon by the Committee shall have lapsed.

         7.4 NO LOANS. Neither the Corporation, any company with which it is
affiliated, nor any of their respective subsidiaries may directly or indirectly
lend money to any person for the purpose of assisting such person in acquiring
or carrying shares of Stock issued upon the exercise of an Option.

         7.5 CONDITIONS PRECEDENT TO EXERCISE. Notwithstanding any other
provision of the Plan, but subject to the provisions of Section 11, the exercise
of an Option following termination of employment or service shall be subject to
the satisfaction of the conditions precedent that the Optionee has not (a)
rendered services or engaged directly or indirectly in any business which in the
opinion of the Committee competes with or is in conflict with the interests of
the Corporation; PROVIDED, HOWEVER, that the ownership by an Optionee of 5% or
less of any class of securities of a publicly traded company shall not be deemed
to violate this clause or (b) violated any written agreement with the
Corporation, including, without limitation, any confidentiality agreement. An
Optionee's violation of clause (a) or (b) of the preceding sentence shall result
in the immediate forfeiture of any Options held by such Optionee.

         7.6 LIMITATIONS ON THE GRANT OF OPTIONS. No one individual may be
granted an Option or Options under the Plan during any fiscal year of the
Corporation for an aggregate number of shares of Stock which exceeds 10% of the
total number of shares reserved for issuance under the Plan; PROVIDED, HOWEVER,
that during the fiscal year of the Corporation ending June 30, 1999 the
Committee may grant Options to the President of the Celera Genomics Group
representing up to 30% of the total number of shares reserved for issuance under
the Plan. The aggregate Fair Market Value of the Stock (determined as of the
date the Option is granted)with respect to which Incentive Stock Options granted
under the Plan and all other stock option plans of the Corporation (or any
parent or subsidiary of the Corporation) are exercisable for the first time by
any specific individual during any calendar year shall not exceed $100,000. No
Incentive Stock Option may be granted hereunder to an individual who immediately
after such Option is granted is a Ten Percent Stockholder unless (a) the Option
price is at least 110% of the


                                      -7-


fair market value of such stock on the date of grant and (b) the Option may not
be exercised more than five (5) years after the date of grant.


8.  EMPLOYEE STOCK AWARDS.

         8.1 GRANT OF EMPLOYEE STOCK AWARDS. Subject to the terms of the Plan,
the Committee may grant Employee Stock Awards to such employees at such time or
times and in such amounts as it shall determine. Shares of Stock issued pursuant
to Employee Stock Awards may, but need not, be subject to such restrictions as
may be established by the Committee at the time of the grant and reflected in an
Agreement.

         8.2 RESTRICTIONS ON EMPLOYEE STOCK AWARDS. Except as provided in the
Plan, any shares of Stock subject to an Employee Stock Award with respect to
which Stock Restrictions have not been satisfied shall be forfeited and all
rights of the employee to such Employee Stock Award shall terminate without any
payment of consideration by the Corporation. Except as set forth in Section 8.5,
a recipient of an Employee Stock Award subject to Stock Restrictions shall
forfeit such award in the event of the termination of his or her employment
during the period the shares are subject to Stock Restrictions.

         8.3 STOCKHOLDER RIGHTS. The recipient of an Employee Stock Award shall
be entitled to such rights of a stockholder with respect to the shares of Stock
issued pursuant to such Employee Stock Award as the Committee shall determine,
including the right to vote such shares of Stock, except that cash and stock
dividends with respect to such shares may, at the discretion of the Committee,
be either paid currently or withheld by the Corporation for the Award
Recipient's account, and interest may be accrued on the amount of cash dividends
withheld at a rate and subject to such terms as determined by the Committee.

         The Committee, in its discretion, may cause a legend or legends to be
placed on any certificate representing shares issued pursuant to Employee Stock
Awards, which legend or legends shall make appropriate reference to the Stock
Restrictions imposed thereon. The Committee may also in its discretion require
that certificates representing shares issued pursuant to Employee Stock Awards
remain in the physical custody of the Corporation or an escrow holder until any
or all of the Stock Restrictions imposed under the Plan have lapsed.

         8.4 NON-TRANSFERABILITY. Prior to the time Stock Restrictions lapse,
none of the shares of Stock issued pursuant to an Employee Stock Award may be
sold, assigned, bequeathed, transferred, pledged, hypothecated, or otherwise
disposed of in any way by the Award Recipient.

         8.5 LAPSE OF RESTRICTIONS. In the event of the termination of
employment of an Award Recipient, prior to the lapse of Stock Restrictions, by
reason of death, total and


                                      -8-



permanent disability, retirement, or resignation or discharge from employment
other than discharge for cause, the Committee may, in its discretion, remove any
Stock Restrictions on all or a portion of the Stock subject to an Employee Stock
Award.

         8.6 LIMITATIONS ON EMPLOYEE STOCK AWARDS. No employee may receive an
Employee Stock Award representing more than 80,000 shares of Stock during any
fiscal year of the Corporation, and the maximum number of shares of Stock which
may be issued to all employees pursuant to Employee Stock Awards under the Plan
shall be 160,000, subject in each case to adjustment in accordance with Section
15.

9.  PERFORMANCE SHARE AWARDS.

         9.1 GRANT OF PERFORMANCE SHARE AWARDS. Subject to the terms of the
Plan, the Committee may grant Performance Share Awards to such employees at such
time or times and in such amounts as it shall determine. Stock issued pursuant
to a Performance Share Award shall be subject to the attainment of performance
goals relating to one or more criteria within the meaning of Section 162(m) of
the Code and the Treasury Regulations issued pursuant thereto, including,
without limitation, stock price, market share, sales, earnings per share, return
on equity, costs, and cash flow, as determined by the Committee from time to
time. Any such objectives and the period in which such objectives are to be met
shall be determined by the Committee at the time of the grant and reflected in
an Agreement; PROVIDED, HOWEVER, that the period in which such objectives are to
be met shall be not less than one year. Each Performance Share Award shall also
be subject to such other restrictions as the Committee may determine.

         9.2 DELIVERY OF PERFORMANCE SHARES. Certificates representing
Performance Shares shall be registered in the Award Recipient's name but shall
remain in the physical custody of the Corporation until the Committee has
determined that the performance goals and other Stock Restrictions with respect
to such Performance Shares have been met.

         9.3 STOCKHOLDER RIGHTS. The recipient of a Performance Share Award
shall be entitled to such rights of a stockholder with respect to the
Performance Shares as the Committee shall determine, including the right to vote
such shares of Stock, except that cash and stock dividends with respect to the
Performance Shares may, at the discretion of the Committee, be either paid
currently or withheld by the Corporation for the Award Recipient's account, and
interest maybe accrued on the amount of cash dividends withheld at a rate and
subject to such terms as determined by the Committee.

         9.4 NON-TRANSFERABILITY. Prior to the time shares of Stock issued
pursuant to a Performance Share Award are delivered to an Award Recipient, none
of such shares may be sold, assigned, bequeathed, transferred, pledged,
hypothecated, or otherwise disposed of in any way by the Award Recipient.

         9.5 LAPSE OF RESTRICTIONS. In the event of the termination of
employment of an Award Recipient, prior to the lapse of Stock Restrictions, by
reason of death, total and


                                      -9-



permanent disability, retirement, or resignation or discharge from employment
other than discharge for cause, the Committee may, in its discretion, remove any
Stock Restrictions on all or a portion of a Performance Share Award, or
determine the performance objectives with respect to all or a portion of a
Performance Share Award to have been attained; PROVIDED, HOWEVER, that the
Committee shall not be entitled to exercise such discretion to the extent that
the ability to exercise such discretion would cause the Performance Share Award
not to qualify as performance based compensation under Section 162(m) of the
Code.

         9.6 LIMITATIONS ON PERFORMANCE SHARE AWARDS. No employee may receive
Performance Share Awards representing more than 200,000 shares of Stock during
any fiscal year of the Corporation, and the maximum number of shares of Stock
which may be issued to all employees pursuant to Performance Share Awards under
the Plan shall be 800,000, subject in each case to adjustment in accordance with
Section 15.

10.  DIRECTOR STOCK AWARDS.

         10.1 GRANT OF DIRECTOR STOCK AWARDS. As of the date of each election or
reelection to the Board of Directors, each Non-Employee Director shall
automatically be granted a Director Stock Award with respect to 300 shares of
Stock, subject to adjustment in accordance with Section 15. Notwithstanding the
foregoing, each Non-Employee Director first elected to the Board of Directors on
a date other than the date of an annual meeting of stockholders shall be granted
that number of whole shares of Stock equal to the number of shares then subject
to a Director Stock Award (after giving effect to Section 10.8) multiplied by a
fraction, the numerator of which shall be the number of months remaining until
the anticipated date of the next annual meeting of stockholders, and the
denominator of which shall be 12. All Director Stock Awards shall be evidenced
by an agreement containing such terms and conditions consistent with the Plan as
the Committee shall determine.

         10.2 VESTING. Each Director Stock Award shall vest in full on the date
immediately preceding the first annual meeting of stockholders next following
the date of grant; PROVIDED, HOWEVER, that, except as provided in the Plan, the
recipient thereof continues to serve as a member of the Board of Directors as of
such date.

         10.3 FORFEITURE OF DIRECTOR STOCK AWARDS. Except as provided in the
Plan, a recipient of a Director Stock Award shall forfeit any unvested shares of
Stock subject to the Director Stock Award, and all rights of the Non-Employee
Director to such unvested shares shall terminate without payment of
consideration by the Corporation, upon the termination of his or her service as
a member of the Board of Directors.

         10.4 STOCKHOLDER RIGHTS. Except as provided in Sections 10.5 and 10.7,
a recipient of a Director Stock Award shall be entitled to all rights of a
stockholder with respect to the shares of Stock issued pursuant to the Director
Stock Award, including the right to receive dividends and to vote such shares of
Stock; PROVIDED, HOWEVER, that stock


                                      -10-


dividends paid with respect to such shares shall be restricted to the same
extent as the underlying shares of Stock issued pursuant to the Director Stock
Award.

         The Committee shall cause a legend or legends to be placed on any
certificate representing shares issued pursuant to a Director Stock Award, which
legend or legends shall make appropriate reference to the terms of the Director
Stock Award and the Plan. The Committee shall also require that certificates
representing shares issued pursuant to Director Stock Awards remain in the
physical custody of the Corporation or an escrow holder until such shares have
vested in accordance with the terms of the Plan.

         10.5 NON-TRANSFERABILITY. Prior to vesting, none of the shares of Stock
issued pursuant to a Director Stock Award may be sold, assigned, bequeathed,
transferred, pledged, hypothecated, or otherwise disposed of in any way by the
recipient thereof.

         10.6 TERMINATION OF SERVICE. If a Non-Employee Director to whom a
Director Stock Award has been granted shall cease to serve as a director as a
result of (a) his or her death, (b) retiring from the Board of Directors upon
reaching Normal Retirement Age, (c) becoming totally and permanently disabled,
or (d) resigning with the approval of the Board of Directors, all shares subject
to such Director Stock Award shall be vested in full, notwithstanding the
provisions of Section 10.2, as of the date of termination of service.

         10.7 DEFERRAL ELECTION. A Non-Employee Director may elect to defer
receipt of any Director Stock Award by filing the appropriate deferral form with
the Corporate Secretary on or before December 15th of the calendar year prior to
the calendar year in which such deferral is to be effective. Notwithstanding the
foregoing, any person elected as a Non-Employee Director for the first time
shall be permitted to make his or her first deferral election no later than
twenty (20) days after such election. In no event, however, shall any deferral
be permitted to the extent prohibited by applicable law. Deferrals shall be
subject to the following terms and conditions:

                  (a) A Non-Employee Director may elect to defer receipt of a
         Director Stock Award until (i) a specified date in the future, (ii)
         cessation of his or her service as a member of the Board of Directors,
         or (iii) the end of the calendar year in which cessation of his or her
         service as a member of the Board of Directors occurs.

                  (b) There shall be established a Deferral Account on the books
         of the Corporation for each Non-Employee Director electing to defer a
         Director Stock Award pursuant to this Section 10.7. Deferrals shall be
         credited to the Non-Employee Director's Deferral Account in Stock Units
         in the following manner: on the award date to which the deferral
         election applies, the amount deferred shall be converted into a number
         of Stock Units equal to the number of shares of Stock awarded that are
         subject to the deferral election. A Non-Employee Director shall not
         have any voting rights with respect to any Stock Units held in his or
         her Deferral Account.


                                      -11-


                  (c) Whenever cash dividends are paid with respect to shares of
         Stock, each Non-Employee Director's Deferral Account shall be credited
         on the payment date of such dividend with additional Stock Units
         (including fractional units to the nearest one/one hundredth (1/100))
         equal in value to the amount of the cash dividend paid on a single
         share of Stock multiplied by the number of Stock Units (including
         fractional units) credited to his or her Deferral Account as of the
         date of record for dividend purposes. For purposes of crediting
         dividends, the value of a Stock Unit shall be the Fair Market Value of
         a share of Stock as of the payment date of the dividend.

                  (d) The number of Stock Units credited to each Non-Employee
         Director's Deferral Account shall be appropriately adjusted in the same
         manner and to the same extent Director Stock Awards are adjusted and
         modified pursuant to Section 15. In the event of a transaction subject
         to Section 11, the Board of Directors shall have the authority to amend
         the Plan to provide for the conversion of Stock Units credited to
         Deferral Accounts into units equal to shares of stock of the resulting
         or acquiring company (or a related company), as appropriate, if such
         stock is publicly traded or, if not, into cash of equal value on the
         effective date of such transaction. If pursuant to the preceding
         sentence cash is credited to a Non-Employee Director's Deferral
         Account, interest shall be credited thereon from the date such cash is
         received to the date of distribution quarterly, at the end of each
         calendar quarter, at a rate per annum (computed on the basis of a
         360-day year and a 91-day quarter) equal to the prime rate announced
         publicly by Citibank, N.A. at the end of such calendar quarter. If
         units representing publicly traded stock of the resulting or acquired
         company (or a related company) are credited to a Non-Employee
         Director's Deferral Account, dividends shall be credited thereto in the
         same manner as dividends are credited on Stock Units credited to such
         Deferral Accounts.

                  (e) Subject to Section 10.7(g), distributions of a
         Non-Employee Director's Deferral Account under the Plan shall be made
         as follows:

                           (i) If a Non-Employee Director has elected to defer a
                  Director Stock Award to a specified date in the future,
                  payment shall be as of such date and shall be made or shall
                  commence, as the case may be, within thirty (30) days after
                  the date specified;

                           (ii) If a Non-Employee Director has elected to defer
                  a Director Stock Award until cessation of his or her service
                  as a member of the Board of Directors, payment shall be as of
                  the date of such cessation of service and shall be made or
                  shall commence, as the case may be, within thirty (30) days
                  after the cessation of the Non-Employee Director's service as
                  a director; and


                                      -12-


                           (iii) If a Non-Employee Director has elected to defer
                  a Director Stock Award until the end of the calendar year in
                  which the cessation of his or her service as a member of the
                  Board of Directors occurs, payment shall be made as of
                  December 31st of such year and shall be made or commence, as
                  the case may be, on December 31st of such year.

                  (f) Notwithstanding any elections pursuant to Sections 10.7(a)
         and/or (g) hereof, in the event of the death of the Non-Employee
         Director prior to the distribution of his or her Deferral Account, the
         balance credited to such Deferral Account as of the date of his or her
         death shall be paid, as soon as reasonably possible thereafter, in a
         single distribution to the Non-Employee Director's beneficiary or
         beneficiaries designated on such Non-Employee Director's deferral
         election form. If no such election or designation has been made, such
         amounts shall be payable to the Non-Employee Director's estate.

                  (g) A Non-Employee Director may elect to have his or her
         Deferral Account under the Plan paid in a single distribution or equal
         annual installments, not to exceed ten (10) annual installments. To the
         extent a Deferral Account is deemed invested in Stock Units, such Stock
         Units shall be converted to Stock on the distribution date as provided
         in Section 10.7(h). To the extent deemed invested in units of any other
         stock, such units shall similarly be converted and distributed in the
         form of stock. To the extent invested in a medium other than Stock
         Units or other units, each such distribution hereunder shall be in the
         medium credited to the Deferral Account.

                  (h) To the extent a Deferral Account is deemed invested in
         Stock Units, a single distribution shall consist of the number of whole
         shares of Stock equal to the number of Stock Units credited to the
         Non-Employee Director's Deferral Account on the date as of which the
         distribution occurs. Cash shall be paid to a Non-Employee Director in
         lieu of a fractional share, determined by reference to the Fair Market
         Value of a share of Stock on the date as of which the distribution
         occurs. In the event a Non-Employee Director has elected to receive
         annual installment payments, each such payment shall be determined as
         follows:

                           (i) To the extent his or her Deferral Account is
                  deemed to be invested in Stock Units, each such payment shall
                  consist of the number of whole shares of Stock equal to the
                  number of Stock Units (including fractional units) credited to
                  the Deferral Account on the date as of which the distribution
                  occurs, divided by the number of annual installments remaining
                  as of such distribution date. Cash shall be paid to
                  Non-Employee Directors in lieu of fractional shares,
                  determined by reference to the Fair Market Value of a share of
                  Stock on the date as of which the distribution occurs.

                           (ii) To the extent his or her Deferral Account has
                  been credited in cash, each such payment shall be calculated
                  by dividing the value on the



                                      -13-


                  date the distribution occurs of that portion of the
                  Non-Employee Director's Deferral Account which is in cash by
                  the number of annual installments remaining as of such
                  distribution date.

         10.8 LIMITATIONS ON DIRECTOR STOCK AWARDS. The Aggregate Value (as such
term is defined below) of all Director Stock Awards granted to a Non-Employee
Director under the Plan in any fiscal year, together with all Director Stock
Awards granted to such Non-Employee Director under the Applera
Corporation/Applied Biosystems Group 1999 Stock Incentive Plan during such year,
may not exceed the Director Budgeted Amount (as such term is defined below) for
such year. In the event that the Aggregate Value of such awards exceeds the
Director Budgeted Amount, the Committee shall allocate to each Non-Employee
Director shares of Stock and shares of Applera Corporation - Applied Biosystems
Group Common Stock ("Applied Biosystems Stock"), in the ratio, as near as may be
practicable, of the number of shares of Stock then outstanding to the number of
shares of Applied Biosystems Stock then outstanding, as determined by the
Committee. For purposes of the Plan, "Aggregate Value" shall be calculated as
the Fair Market Value of a share of Stock or Applied Biosystems Stock, as the
case may be, on the date of grant multiplied by the number of shares of Stock or
Applied Biosystems Stock, as the case may be, subject to the Director Stock
Award, and the "Director Budgeted Amount" shall be $135,000 for the fiscal year
ending June 30, 2001 and shall increase by 10% for each fiscal year thereafter.

11.  ACCELERATION UPON A CHANGE OF CONTROL.

         Notwithstanding any other provision of the Plan or any Option or Award
granted hereunder, (a) any Option granted hereunder and then outstanding shall
become immediately exercisable in full, (b) all Stock Restrictions shall
immediately terminate, and (c) all performance objectives applicable to any
Performance Share Award shall be deemed attained (i) in the event that a tender
offer or exchange offer (other than an offer by the Corporation) for common
stock of the Company representing more than 25% of the combined voting power of
the then outstanding voting securities of the Company entitled to vote generally
in the election of directors ("Voting Securities") is made by any "person"
within the meaning of Section 14(d) of the Act and not withdrawn within ten (10)
days after the commencement thereof; PROVIDED, HOWEVER, that the Committee may
by action taken prior to the end of such ten (10) day period extend such ten
(10) day period; and, PROVIDED FURTHER, that the Committee may by further action
taken prior to the end of such extended period declare (a) all Options granted
hereunder and then outstanding to be immediately exercisable in full, (b) all
Stock Restrictions to be immediately terminated, and (c) all performance
objectives applicable to any Performance Share Award to be deemed attained; or
(ii) in the event of a Change in Control (as hereinafter defined).

         For purposes of this Section 11, a "Change in Control" means an event
that would be required to be reported (assuming such event has not been
"previously reported") in response to Item 1(a) of the Current Report on Form
8-K, as in effect on the effective date of the Plan, pursuant to Section 13 or
15(d) of the Act; PROVIDED, HOWEVER, that,


                                      -14-



without limitation, such a Change in Control shall be deemed to have occurred at
such time as (a) any "person" within the meaning of Section 14(d) of the Act
becomes the "beneficial owner" as defined in Rule 13d-3 thereunder, directly or
indirectly, of more than 25% of the combined voting power of the then
outstanding Voting Securities, (b) during any two-year period, individuals who
constitute the Board of Directors (the "Incumbent Board") as of the beginning of
the period cease for any reason to constitute at least a majority thereof,
provided that any person becoming a director during such period whose election
or nomination for election by the Corporation's stockholders was approved by a
vote of at least three-quarters of the Incumbent Board (either by a specific
vote or by approval of the proxy statement of the Corporation in which such
person is named as a nominee for director without objection to such nomination,
other than in response to an actual or threatened Change in Control or proxy
contest) shall be, for purposes of this clause (b), considered as though such
person were a member of the Incumbent Board, or (c) the approval by the
Corporation's stockholders of the sale of all or substantially all of the stock
or assets of the Corporation. The Committee may adopt such procedures as to
notice and exercise as may be necessary to effectuate the acceleration of the
exercisability of Options, termination of Stock Restrictions, and attainment of
performance objectives as described above.

12.  SHARE WITHHOLDING.

         With respect to any Option or Award, the Committee may, in its
discretion and subject to such rules as the Committee may adopt, permit or
require any Optionee or Award Recipient to satisfy, in whole or in part, any
withholding tax obligation which may arise in connection with an Option or Award
by electing to have the Corporation withhold Stock having a Fair Market Value
(as of the date the amount of withholding tax is determined) equal to the amount
of withholding tax.

13.  NO RIGHT TO CONTINUED EMPLOYMENT OR SERVICE.

         Nothing contained in the Plan or in any Option or Award granted or
Agreement entered into pursuant to the Plan shall confer upon any employee the
right to continue in the employ of the Corporation, any consultant the right to
continue to perform services for the Corporation, or any Non-Employee Director
the right to continue as a member of the Board of Directors or interfere with
the right of the Corporation to terminate such employee's employment, such
consultant's service, or Non-Employee Director's service at any time.

14.  TIME OF GRANTING OPTIONS AND EMPLOYEE AWARDS.

         Nothing contained in the Plan or in any resolution adopted by the Board
of Directors or the holders of Stock shall constitute the grant of any Option or
Award hereunder. An Option or Award under the Plan shall be deemed to have been
granted on the date on which the name of the recipient and the terms of the
Option or Award are set forth in an Agreement and delivered to the recipient,
unless otherwise provided in the Agreement.


                                      -15-



15.  ADJUSTMENTS UPON CHANGES IN CAPITALIZATION.

         Notwithstanding any other provision of the Plan, in the event of
changes in the outstanding Stock by reason of stock dividends, stock splits,
recapitalizations, combinations or exchanges of shares, corporate separations or
divisions (including, but not limited to, split-ups, split-offs, or spin-offs),
reorganizations (including, but not limited to, mergers or
consolidations),liquidations, or other similar events, the aggregate number and
class of shares available under the Plan, the number of shares subject to
Director Stock Awards, the maximum number of shares that may be subject to
Options and Awards, and the terms of any outstanding Options or Awards
(including, without limitation, the number of shares subject to an outstanding
Option or Award and the price at which shares of Stock may be issued pursuant to
an outstanding Option) and of any Stock Units shall be adjusted in such manner
as the Committee in its discretion deems appropriate.

16.  TERMINATION AND AMENDMENT OF THE PLAN.

         Unless the Plan shall have been terminated as hereinafter provided, no
Option or Award shall be granted hereunder after March 31, 2004. The Board of
Directors may at any time prior to that date terminate the Plan or make such
modification or amendment to the Plan as it shall deem advisable; PROVIDED,
HOWEVER, that, except as provided in Section 15, no amendment may be made
without the approval by the holders of Stock (to the extent such approval would
be required for an exemption under Section 16(b) of the Act which the Company
wishes to have) if such amendment would (a) increase the aggregate number of
shares of Stock which may be issued under the Plan, (b) materially modify the
requirements as to eligibility for participation in the Plan, or (c) materially
increase the benefits accruing to participants under the Plan. No termination,
modification, or amendment of the Plan may, without the consent of an Optionee
or Award Recipient, adversely affect in any material manner the rights of such
Optionee or Award Recipient under any Option or Award.

17.  AMENDMENT OF OPTIONS AND AWARDS AT THE DISCRETION OF THE COMMITTEE.

         The terms of any outstanding Option or Award may be amended from time
to time by the Committee in its discretion in any manner that it deems
appropriate, including, without limitation, acceleration of the date of exercise
of any Option or Award, termination of Stock Restrictions as to any Award, or
the conversion of an Incentive Stock Option into a Non-Qualified Stock Option;
PROVIDED, HOWEVER, that no such amendment shall adversely affect in any material
manner any right of any Optionee or Award Recipient under the Plan without his
or her consent; and, PROVIDED FURTHER, that the Committee shall not (a) amend
any previously-issued Performance Share Award to the extent that such amendment
would cause such Performance Share Award not to qualify as performance based
compensation under Section 162(m) of the Code or (b) amend any previously-issued
Option to reduce the purchase price thereof whether by modification of


                                      -16-



the Option or by cancellation of the Option in consideration of the immediate
issuance of a replacement Option bearing a reduced purchase price.

18.  GOVERNMENT REGULATIONS.

         The Plan and the grant and exercise of Options and Awards hereunder,
and the obligation of the Corporation to issue, sell, and deliver shares, as
applicable, under such Options and Awards, shall be subject to all applicable
laws, rules, and regulations. Notwithstanding any other provision of the Plan,
transactions under the Plan are intended to comply with the applicable
exemptions under Rule 16b-3 under the Act as to persons subject to the reporting
requirements of Section 16(a) of the Act with respect to shares of Stock, and
Options and Awards under the Plan shall be fashioned and administered in a
manner consistent with the conditions applicable under Rule 16b-3.

19.  OPTIONS AND AWARDS IN FOREIGN COUNTRIES.

         The Committee shall have the authority and discretion to adopt such
modifications, procedures, and subplans as it shall deem necessary or desirable
to comply with the provisions of the laws of foreign countries in which the
Corporation may operate in order to assure the viability of the benefits of the
Options and Awards made to individuals employed in such countries and to meet
the objectives of the Plan.

20.  GOVERNING LAW.

         The Plan shall be construed, regulated, and administered under the
internal laws of the State of Delaware.

21.  STOCKHOLDER APPROVAL.

         The Plan shall become effective upon the date of adoption by the Board
of Directors, subject to approval by the stockholders of the Corporation in
accordance with applicable law. Unless so approved within one (1) year after the
date of the adoption of the Plan by the Board of Directors, the Plan shall not
be effective for any purpose. Prior to approval by the Corporation's
stockholders, the Committee may grant Options and Awards under the terms of the
Plan, but if stockholder approval is not obtained in the specified period, such
Options and Awards shall be of no effect.





                                      -17-


                      2002 ANNUAL MEETING OF STOCKHOLDERS

                                OCTOBER 17, 2002
                                   9:30 A.M.

               YOU MAY VOTE BY TELEPHONE, BY INTERNET, OR BY MAIL
                       (SEE INSTRUCTIONS ON REVERSE SIDE)

                             YOUR VOTE IS IMPORTANT

                                  DETACH HERE


                                     PROXY

                              APPLERA CORPORATION

   THIS PROXY IS SOLICITED ON BEHALF OF THE BOARD OF DIRECTORS FOR THE ANNUAL
                  MEETING OF STOCKHOLDERS ON OCTOBER 17, 2002


The undersigned stockholder(s) of Applera Corporation (the "Company") hereby
appoints TONY L. WHITE, WILLIAM B. SAWCH, and THOMAS P. LIVINGSTON, and each of
them, as proxy or proxies, with power of substitution to vote all shares of
Applera Corporation - Applied Biosystems Group Common Stock and/or Applera
Corporation - Celera Genomics Group Common Stock which the undersigned is
entitled to vote (including shares, if any, held on behalf of the undersigned,
and indicated on the reverse hereof, by EquiServe Trust Company, under the
Company's dividend reinvestment plan and by Mellon Investor Services, L.L.C.
under the Company's employee stock purchase plans) at the 2002 Annual Meeting of
Stockholders and at any adjournment or adjournments thereof, as indicated on the
reverse side hereof, and, in their discretion, upon such other matters as may
properly come before the meeting, all as more fully described in the Proxy
Statement for such Annual Meeting.

THIS PROXY WHEN PROPERLY EXECUTED AND RETURNED WILL BE VOTED AS DIRECTED ON THE
REVERSE SIDE OR, IF NO DIRECTION IS GIVEN, IT WILL BE VOTED FOR THE ELECTION OF
ALL NOMINEES FOR DIRECTOR AND FOR PROPOSALS 2, 3, 4, AND 5.




SEE REVERSE       CONTINUED AND TO BE SIGNED ON REVERSE SIDE        SEE REVERSE
   SIDE                                                                 SIDE



[APPLERA LOGO]

APPLERA CORPORATION

C/O EQUISERVE
P.O. BOX 43068
PROVIDENCE, RI 02940


                                                     
VOTE BY TELEPHONE                                       VOTE BY INTERNET
It's fast, convenient, and immediate!                   It's fast, convenient, and your vote is immediately
Call Toll-Free on a Touch-Tone Phone                    confirmed and posted.
1-877-PRX-VOTE (1-877-779-8683).


FOLLOW THESE FOUR EASY STEPS:                           FOLLOW THESE FOUR EASY STEPS:

1. READ THE ACCOMPANYING PROXY STATEMENT AND            1. READ THE ACCOMPANYING PROXY STATEMENT AND
   PROXY CARD.                                             PROXY CARD.

2. CALL THE TOLL-FREE NUMBER                            2. GO TO THE WEBSITE
   1-877-PRX-VOTE (1-877-779-8683).                        HTTP://WWW.EPROXYVOTE.COM/ABI

3. ENTER YOUR VOTER CONTROL NUMBER LOCATED ON           3. ENTER YOUR VOTER CONTROL NUMBER LOCATED ON
   YOUR PROXY CARD ABOVE YOUR NAME.                        YOUR PROXY CARD ABOVE YOUR NAME.

4. FOLLOW THE INSTRUCTIONS PROVIDED.                    4. FOLLOW THE RECORDED INSTRUCTIONS.

YOUR VOTE IS IMPORTANT!                                 YOUR VOTE IS IMPORTANT!
Call 1-877-PRX-VOTE anytime!                            Go to HTTP://WWW.EPROXYVOTE.COM/ABI anytime!



DO NOT RETURN YOUR PROXY CARD IF YOU ARE VOTING BY TELEPHONE OR
INTERNET


                                  DETACH HERE

/X/     PLEASE MARK
        VOTES AS IN
        THIS EXAMPLE.

          THE BOARD OF DIRECTORS RECOMMENDS A VOTE FOR ALL PROPOSALS.



                                                                                                                 
                                                                                                                 FOR AGAINST ABSTAIN

1. Election of Directors.                                  2. Ratification of the selection of Pricewaterhouse   / /   / /     / /
                                                              Coopers LLP as independent accountants for the
                                                              fiscal year ending June 30, 2003.

NOMINEES: (01) Richard H. Ayers, (02) Jean-Luc Belingard,  3. Approval of amendments to the Applera              / /   / /     / /
(03) Robert H. Hayes, (04) Arnold J. Levine, (05)             Corporation 1999 Employee Stock Purchase Plan.
Theodore E. Martin, (06) Carolyn W. Slayman, (07) Orin R.
Smith, (08) Georges C. St. Laurent, Jr., (09) James R.     4. Approval of amendment to the Applera              / /   / /     / /
Tobin, and (10) Tony L. White                                 Corporation/Applied Biosystems Group 1999 Stock
                                                              Incentive Plan.

     FOR                WITHHELD                           5. Approval of amendment to the Applera              / /   / /     / /
     ALL                FROM ALL                              Corporation/Celera Genomics Group 1999 Stock
   NOMINEES  / /        NOMINEES  / /                         Incentive Plan.


/ /                                                               MARK HERE FOR                MARK HERE IF
   ----------------------------------------                       ADDRESS CHANGE            YOU PLAN TO ATTEND
    For all nominees except as noted above                       AND NOTE AT LEFT  / /        THE MEETING               / /

                                                             MARK HERE TO DISCONTINUE MAILING ANNUAL REPORT ON THIS
                                                                        ACCOUNT (FOR MULTIPLE ACCOUNTS ONLY)            / /

                                                             PLEASE SIGN EXACTLY AS YOUR NAME APPEARS. IF ACTING AS
                                                             ATTORNEY, EXECUTOR, TRUSTEE, OR IN A REPRESENTATIVE
                                                             CAPACITY, SIGN NAME AND TITLE.

Signature: __________________________ Date: ____________ Signature: ___________________________ Date: __________